Nomenclature for Factors of the HLA System

March 2018

Compiled by Steven G. E. Marsh for the WHO Nomenclature Committee for Factors of the HLA System.

Published In:

The following sequences have been submitted to the Nomenclature Committee since the February 2018 nomenclature update and, following agreed policy, have been assigned official allele designations. Full details of all sequences will be published in a forthcoming report.

Below are listed the newly assigned sequences (Table 1) and confirmations of previously reported sequences (Table 2). The accession number of each sequence is given and these can be used to retrieve the sequence files from the EMBL, GenBank or DDBJ data libraries. Although accession numbers have been assigned by the data-libraries and most sequences are already available, there is still the possibility that an author may not yet have allowed the sequence to be released; in such a case you will have to contact the submitting author directly. Additional information pertaining to new sequences is often included in the publications describing these alleles; a listing of recent publications that describe new HLA sequences is given in Table 3.

All new and confirmatory sequences should now be submitted directly to the WHO Nomenclature Committee for Factors of the HLA System via the IMGT/HLA Database using the sequence submission tool provided (2). The IMGT/HLA Database may be accessed via the world wide web at : http://ww.ebi.ac.uk/imgt/hla/.

Table 1: New Sequences


SequenceCell identificationAccession numberSubmitting author
A*01:01:84HG00014033MG724789,Histogenetics, Ossining, USA
A*01:01:84BST13253640MH045578,Francesc Rudilla, Barcelona, Spain
A*01:249HG00013960MG701435,Histogenetics, Ossining, USA
A*01:250NHG00013984,HG00013991,HG00013992,HG00013985MG701459,MG701466,MG701467,MG701460,Histogenetics, Ossining, USA
A*01:251HG00014062MG724818,Histogenetics, Ossining, USA
A*01:25271706162MG954068,Stephane Buhler, Geneva, Switzerland
A*01:253BST1326564MG999587,Francesc Rudilla, Barcelona, Spain
A*02:01:135HG00013633,HG00013965,HG00013969,HG00013956MG603073,MG701440,MG701444,MG701431,Histogenetics, Ossining, USA
A*02:01:136HG00013966MG701441,Histogenetics, Ossining, USA
A*02:01:137HG00013995MG701470,Histogenetics, Ossining, USA
A*02:06:23HG00014040MG724796,Histogenetics, Ossining, USA
A*02:243:03BST1344744MG999588,Francesc Rudilla, Barcelona, Spain
A*02:711HG00013621MG584012,Histogenetics, Ossining, USA
A*02:712HG00013626MG584017,Histogenetics, Ossining, USA
A*02:713HG00013632MG603072,Histogenetics, Ossining, USA
A*02:714HG00013637MG603077,Histogenetics, Ossining, USA
A*02:715NHG00013646MG603086,Histogenetics, Ossining, USA
A*02:716HG00013948MG701423,Histogenetics, Ossining, USA
A*02:717HG00013958MG701433,Histogenetics, Ossining, USA
A*02:718HG00013961MG701436,Histogenetics, Ossining, USA
A*02:719HG00013962,HG00014027MG701437,MG724783,Histogenetics, Ossining, USA
A*02:720HG00013964MG701439,Histogenetics, Ossining, USA
A*02:721HG00014417MG756785,Histogenetics, Ossining, USA
A*02:72271703468MG954062,Stephane Buhler, Geneva, Switzerland
A*02:723JRC-D78-129LC371766,Masahiro Satake, Tokyo, Japan
A*02:724HG00015476MG930530,Histogenetics, Ossining, USA
A*02:725JXZJ201712188001MH054901,Faming Zhu, Hangzhou, China
A*03:01:65HG00013628MG584019,Histogenetics, Ossining, USA
A*03:01:66HG00013647MG603087,Histogenetics, Ossining, USA
A*03:01:67HG00014413MG756781,Histogenetics, Ossining, USA
A*03:291HG00013951,HG00014017MG701426,MG724773,Histogenetics, Ossining, USA
A*03:292HG00013968MG701443,Histogenetics, Ossining, USA
A*03:293HG00014060MG724816,Histogenetics, Ossining, USA
A*11:01:80HG00013644MG603084,Histogenetics, Ossining, USA
A*11:01:81HG00013996MG701471,Histogenetics, Ossining, USA
A*11:275HG00013638MG603078,Histogenetics, Ossining, USA
A*11:276HG00013986MG701461,Histogenetics, Ossining, USA
A*11:277HG00013989MG701464,Histogenetics, Ossining, USA
A*11:278HG00014023MG724779,Histogenetics, Ossining, USA
A*11:278BST13324760MH45580,Francesc Rudilla, Barcelona, Spain
A*11:27971705468MG954065,Stephane Buhler, Geneva, Switzerland
A*11:280JXZJ201712122014MH054902,Faming Zhu, Hangzhou, China
A*23:01:21HG00014001MG701476,Histogenetics, Ossining, USA
A*23:17:01:02HG00013858,HG00013869MG724749,MG724765,Histogenetics, Ossining, USA
A*23:87MGM_131701MG886871,Vibhuti Rambani, Bangalore, India
A*23:88HG00015469,HG00015474MG930523,MG930528,Histogenetics, Ossining, USA
A*24:02:105HG00013629MG584020,Histogenetics, Ossining, USA
A*24:02:106HG00013998MG701473,Histogenetics, Ossining, USA
A*24:398HG00013641MG603081,Histogenetics, Ossining, USA
A*24:399HG00013643MG603083,Histogenetics, Ossining, USA
A*24:400HG00013980MG701455,Histogenetics, Ossining, USA
A*24:401BST010MG967421,Francesc Rudilla, Barcelona, Spain
A*24:402BST1339249MG999589,Francesc Rudilla, Barcelona, Spain
A*26:01:46HG00013853,HG00013857,HG00013864,HG00013868,HG00014031,HG00014043,HG00014044,HG00015475MG701415,MG724748,MG724756,MG724762,MG724787,MG724799,MG724800,MG930529,Histogenetics, Ossining, USA
A*30:02:12HG00005755KC503857,Histogenetics, Ossining, USA
A*30:128HG00014003MG701478,Histogenetics, Ossining, USA
A*30:1293110129MG999642,Antonio Balas, Madrid, Spain
A*31:01:28HG00013854MG701416,Histogenetics, Ossining, USA
A*31:135HG00013645,HG00013945,HG00013953,HG00013976,HG00014016,HG00014056MG603085,MG701420,MG701428,MG701451,MG724772,MG724812,Histogenetics, Ossining, USA
A*32:01:29HG00013954MG701429,Histogenetics, Ossining, USA
A*32:109HG00014416MG756784,Histogenetics, Ossining, USA
A*32:110HG00013974,HG00014038MG701449,MG724794,Histogenetics, Ossining, USA
A*33:03:34HG00014002,HG00014004MG701477,MG701479,Histogenetics, Ossining, USA
A*33:142HG00013622MG584013,Histogenetics, Ossining, USA
A*33:142HG00013990MG701465,Histogenetics, Ossining, USA
A*33:143NHG00013631MG603071,Histogenetics, Ossining, USA
A*33:144HG00013639MG603079,Histogenetics, Ossining, USA
A*33:145HG00013978MG701453,Histogenetics, Ossining, USA
A*33:146HG00015470MG930524,Histogenetics, Ossining, USA
A*36:06BST13346840MH045581,Francesc Rudilla, Barcelona, Spain
A*66:31HG00014024MG724780,Histogenetics, Ossining, USA
A*68:01:36HG00013634MG603074,Histogenetics, Ossining, USA
A*68:175HG00013640MG603080,Histogenetics, Ossining, USA
A*68:176HG00014037MG724793,Histogenetics, Ossining, USA
A*68:17771704723MG954064,Stephane Buhler, Geneva, Switzerland
A*74:01:082016-7070MH028637,Mauro Castagnetta, Genova, Italy
B*07:04:02HG00014418MG756787,Histogenetics, Ossining, USA
B*07:06:03HG00013678MG603096,Histogenetics, Ossining, USA
B*07:306HG00013681MG603099,Histogenetics, Ossining, USA
B*07:307HG00013689MG603107,Histogenetics, Ossining, USA
B*07:308HG00014072MG701492,Histogenetics, Ossining, USA
B*07:309HG00014093MG701513,Histogenetics, Ossining, USA
B*07:310HG00014096,HG00014421MG701516,MG756790,Histogenetics, Ossining, USA
B*07:311HG00014112MG724842,Histogenetics, Ossining, USA
B*08:01:38HG00013879MG724825,Histogenetics, Ossining, USA
B*08:01:39HG00014454MG756826,Histogenetics, Ossining, USA
B*08:191HG00013648MG584021,Histogenetics, Ossining, USA
B*08:192HG00013670MG603088,Histogenetics, Ossining, USA
B*08:193HG00013690,HG00013691MG603108,MG603109,Histogenetics, Ossining, USA
B*08:194HG00014436MG756805,Histogenetics, Ossining, USA
B*08:1958109342MG999640,Antonio Balas, Madrid, Spain
B*13:01:13HG00013675,HG00013687,HG00013688MG603093,MG603105,MG603106,Histogenetics, Ossining, USA
B*13:02:22HG00013673MG603091,Histogenetics, Ossining, USA
B*13:110327945MG972369,Elena Longhi, Milano, Italy
B*14:61HG00014071MG701491,Histogenetics, Ossining, USA
B*15:03:07HG00014445MG756814,Histogenetics, Ossining, USA
B*15:18:01:04CNUHH171204_71855689MG992019,Oh-Joong Kwon, Seoul, South Korea
B*15:440HG00013669MG584042,Histogenetics, Ossining, USA
B*15:441HG00014074MG701494,Histogenetics, Ossining, USA
B*15:442HG00014094MG701514,Histogenetics, Ossining, USA
B*15:443HG00014107MG724837,Histogenetics, Ossining, USA
B*15:444HG00014430MG756799,Histogenetics, Ossining, USA
B*15:445HG00014442MG756811,Histogenetics, Ossining, USA
B*18:14446721LT898191,Marcel GJ Tilanus, Maastricht, The Netherlands
B*18:145HG00013873,HG00013878,HG00013881,HG00013889,HG00014075,HG00014110,HG00014113MG701483,MG724824,MG724828,MG724836,MG701495,MG724840,MG724843,Histogenetics, Ossining, USA
B*27:05:33HG00014447MG756818,Histogenetics, Ossining, USA
B*27:173HG00013684MG603102,Histogenetics, Ossining, USA
B*27:174HG00013871MG701481,Histogenetics, Ossining, USA
B*27:175BST009MG967420,Francesc Rudilla, Barcelona, Spain
B*35:01:48HG00014086MG701506,Histogenetics, Ossining, USA
B*35:364HG00013649MG584022,Histogenetics, Ossining, USA
B*35:365HG00014077,HG00014068,HG00014070MG701497,MG701480,MG701488,MG701490,Histogenetics, Ossining, USA
B*35:366HG00014091MG701511,Histogenetics, Ossining, USA
B*35:367HG00014103MG701523,Histogenetics, Ossining, USA
B*35:368HG00014453MG756825,Histogenetics, Ossining, USA
B*35:369BST1331479MG925815,Francesc Rudilla, Barcelona, Spain
B*35:3702017-2415MG999575,Mauro Castagnetta, Genova, Italy
B*38:69HG00013692MG603110,Histogenetics, Ossining, USA
B*38:70HG00014441MG756810,Histogenetics, Ossining, USA
B*39:05:03HG00014114,HG00014419,HG00014443,HG00013877,HG00014069,HG00013656MG724844,MG756788,MG756812,MG724822,MG701489,MG584029,Histogenetics, Ossining, USA
B*39:134328667MG972370,Elena Longhi, Milano, Italy
B*40:01:50HG00013693MG603111,Histogenetics, Ossining, USA
B*40:01:5117ZZ0727LC371656,Li-Qun Zhang, Beijing, China
B*40:01:52BST1316456MH022878,Francesc Rudilla, Barcelona, Spain
B*40:367HG00013680MG603098,Histogenetics, Ossining, USA
B*40:368HG00014104MG701524,Histogenetics, Ossining, USA
B*40:369HG00014431MG756800,Histogenetics, Ossining, USA
B*40:370HG00014446MG756815,Histogenetics, Ossining, USA
B*42:25HG00013650,HG00013887,HG00014423MG584023,MG724834,MG756792,Histogenetics, Ossining, USA
B*44:02:49HG00013672,HG00013882MG603090,MG724829,Histogenetics, Ossining, USA
B*44:27:03HG00014451MG756823,Histogenetics, Ossining, USA
B*44:163:02HG00013674MG603092,Histogenetics, Ossining, USA
B*44:279HG00013653MG584026,Histogenetics, Ossining, USA
B*44:280HG00013655MG584028,Histogenetics, Ossining, USA
B*44:281HG00014440MG756809,Histogenetics, Ossining, USA
B*44:282HG00014448MG756819,Histogenetics, Ossining, USA
B*44:283JRC-D08-101LC371763,Masahiro Satake, Tokyo, Japan
B*48:44HG00013671MG603089,Histogenetics, Ossining, USA
B*48:45HG00014428MG756797,Histogenetics, Ossining, USA
B*48:46JRC-D78-156LC371765,Masahiro Satake, Tokyo, Japan
B*49:49HG00013667MG584040,Histogenetics, Ossining, USA
B*49:50HG00014452MG756824,Histogenetics, Ossining, USA
B*51:01:59HG00013652MG584025,Histogenetics, Ossining, USA
B*51:01:60HG00013875,HG00013888,HG00014422MG701485,MG724835,MG756791,Histogenetics, Ossining, USA
B*51:01:61HG00014108MG724838,Histogenetics, Ossining, USA
B*51:01:621024563MG99941,Antonio Balas, Madrid, Spain
B*51:06:04HG00014090MG701510,Histogenetics, Ossining, USA
B*51:232HG00014079MG701499,Histogenetics, Ossining, USA
B*51:233HG00014427MG756796,Histogenetics, Ossining, USA
B*51:234HG00014433,HG00014434MG756802,MG756803,Histogenetics, Ossining, USA
B*52:01:30HG00014080MG701500,Histogenetics, Ossining, USA
B*52:09:02JRC-D36-077LC371764,Masahiro Satake, Tokyo, Japan
B*52:74HG00013679,HG00013686MG603097,MG603104,Histogenetics, Ossining, USA
B*52:75HG00014087MG701507,Histogenetics, Ossining, USA
B*53:01:15HG00014116MG724846,Histogenetics, Ossining, USA
B*53:50HG00014099MG701519,Histogenetics, Ossining, USA
B*56:01:10HG00013880MG724827,Histogenetics, Ossining, USA
B*56:01:11HG00014089MG701509,Histogenetics, Ossining, USA
B*56:04:02HG00014420MG756789,Histogenetics, Ossining, USA
B*57:01:25HG00013694MG603112,Histogenetics, Ossining, USA
B*57:01:26HG00013872MG701482,Histogenetics, Ossining, USA
B*57:01:27HG00014435MG756804,Histogenetics, Ossining, USA
B*57:04:03HG00014098MG701518,Histogenetics, Ossining, USA
B*57:94HG00013651MG584024,Histogenetics, Ossining, USA
B*57:95HG00014088MG701508,Histogenetics, Ossining, USA
B*57:96HG00014450MG756821,Histogenetics, Ossining, USA
B*58:01:23HG00013658MG584031,Histogenetics, Ossining, USA
B*58:93NHG00013666MG584039,Histogenetics, Ossining, USA
C*01:02:44HG00013738MG603134,Histogenetics, Ossining, USA
C*01:02:45HG00014469MG756843,Histogenetics, Ossining, USA
C*01:146:02HG00014466MG756839,Histogenetics, Ossining, USA
C*01:146:02DKMS-LSL-C-2397LT989985,DKMS Life Sciences Lab, Dresden, Germany
C*01:149HG00013730MG603126,Histogenetics, Ossining, USA
C*01:150HG00014131MG701543,Histogenetics, Ossining, USA
C*01:151HG00014144,HG00014198MG701556,MG724897,Histogenetics, Ossining, USA
C*01:152HG00014152MG701564,Histogenetics, Ossining, USA
C*01:153HG00014491MG756865,Histogenetics, Ossining, USA
C*02:02:35HG00014154MG701566,Histogenetics, Ossining, USA
C*02:02:36HG00014195MG724894,Histogenetics, Ossining, USA
C*02:02:37BST011MH006687,Francesc Rudilla, Barcelona, Spain
C*02:136HG00014455MG756827,Histogenetics, Ossining, USA
C*02:137NT00671DQ648007,Carolyn K Hurley, Washington DC, USA
C*02:137NT02062MG952918,Carolyn K Hurley, Washington DC, USA
C*02:138M2MG976799,Brian Dukek, Rochester, USA
C*02:139DKMS-LSL-C-2390LT989978,DKMS Life Sciences Lab, Dresden, Germany
C*03:03:38DKMS-LSL-C-2403LT989991,DKMS Life Sciences Lab, Dresden, Germany
C*03:03:39HG00014480MG756854,Histogenetics, Ossining, USA
C*03:03:40DKMS-LSL-C-2386LT989974,DKMS Life Sciences Lab, Dresden, Germany
C*03:04:58HG00014141MG701553,Histogenetics, Ossining, USA
C*03:296:02HG00013709,HG00013707MG584057,MG584055,Histogenetics, Ossining, USA
C*03:378HG00013905MG724861,Histogenetics, Ossining, USA
C*03:379HG00014128,HG00014487MG701540,MG756861,Histogenetics, Ossining, USA
C*03:380NHG00014137,HG00014456MG701549,MG756828,Histogenetics, Ossining, USA
C*03:381HG00014143MG701555,Histogenetics, Ossining, USA
C*03:382HG00014158MG701570,Histogenetics, Ossining, USA
C*03:383BST1346518MG976847,Jose Luis Caro Oleas, Barcelona, Spain
C*03:384JRC-D78-128LC371761,Masahiro Satake, Tokyo, Japan
C*03:385BST1307387MG820615,Francesc Rudilla, Barcelona, Spain
C*03:386BST1322338MH022881,Francesc Rudilla, Barcelona, Spain
C*03:387BST1322036MH022880,Francesc Rudilla, Barcelona, Spain
C*03:388DKMS-LSL-C-2400LT989988,DKMS Life Sciences Lab, Dresden, Germany
C*04:01:86HG00014127MG701539,Histogenetics, Ossining, USA
C*04:01:87HG00014157MG701569,Histogenetics, Ossining, USA
C*04:01:88DKMS-LSL-C-2391LT989979,DKMS Life Sciences Lab, Dresden, Germany
C*04:01:89DKMS-LSL-C-2389LT989977,DKMS Life Sciences Lab, Dresden, Germany
C*04:01:90DKMS-LSL-C-2433LT990021,DKMS Life Sciences Lab, Dresden, Germany
C*04:01:91DKMS-LSL-C-2428LT990016,DKMS Life Sciences Lab, Dresden, Germany
C*04:201:02JRC-D08-094LC371759,Masahiro Satake, Tokyo, Japan
C*04:289HG00013899,HG00013902MG724855,MG724858,Histogenetics, Ossining, USA
C*04:290HG00014150MG701562,Histogenetics, Ossining, USA
C*04:291DKMS-LSL-C-2415LT990003,DKMS Life Sciences Lab, Dresden, Germany
C*04:292DKMS-LSL-C-2396LT989984,DKMS Life Sciences Lab, Dresden, Germany
C*04:293DKMS-LSL-C-2434LT990022,DKMS Life Sciences Lab, Dresden, Germany
C*05:01:37HG00014130MG701542,Histogenetics, Ossining, USA
C*05:01:38HG00014465MG756838,Histogenetics, Ossining, USA
C*05:159HG00013741MG603137,Histogenetics, Ossining, USA
C*05:159DKMS-LSL-C-2410LT989998,DKMS Life Sciences Lab, Dresden, Germany
C*05:160HG00014498MG756873,Histogenetics, Ossining, USA
C*05:161BST1316803MG820617,Francesc Rudilla, Barcelona, Spain
C*05:162DKMS-LSL-C-2405LT989993,DKMS Life Sciences Lab, Dresden, Germany
C*05:163DKMS-LSL-C-2439LT990027,DKMS Life Sciences Lab, Dresden, Germany
C*05:164DKMS-LSL-C-2448LT990036,DKMS Life Sciences Lab, Dresden, Germany
C*06:02:01:12NT2085AMG986061,Carolyn K Hurley, Washington DC, USA
C*06:02:50:02HG00014138MG701550,Histogenetics, Ossining, USA
C*06:02:51HG00013695MG584043,Histogenetics, Ossining, USA
C*06:02:52DKMS-LSL-C-2395LT989983,DKMS Life Sciences Lab, Dresden, Germany
C*06:213HG00014129MG701541,Histogenetics, Ossining, USA
C*06:21471705092MG954066,Stephane Buhler, Geneva, Switzerland
C*06:215N60785MH031344,Runying Tian, Durham, USA
C*06:216DKMS-LSL-C-2413LT990001,DKMS Life Sciences Lab, Dresden, Germany
C*06:217DKMS-LSL-C-2398LT989986,DKMS Life Sciences Lab, Dresden, Germany
C*06:218DKMS-LSL-C-2408LT989996,DKMS Life Sciences Lab, Dresden, Germany
C*07:01:60HG00013712MG584060,Histogenetics, Ossining, USA
C*07:01:61HG00014146MG701558,Histogenetics, Ossining, USA
C*07:02:84HG00014160MG701572,Histogenetics, Ossining, USA
C*07:02:85HG00014184,HG00014478,HG00014134,HG00014136,HG00013702MG724883,MG756852,MG701546,MG701548,MG584050,Histogenetics, Ossining, USA
C*07:02:86DKMS-LSL-C-2394LT989982,DKMS Life Sciences Lab, Dresden, Germany
C*07:04:12DKMS-LSL-C-2411LT989999,DKMS Life Sciences Lab, Dresden, Germany
C*07:73:02HG00013735MG603131,Histogenetics, Ossining, USA
C*07:614HG00013713MG584061,Histogenetics, Ossining, USA
C*07:615HG00013717MG603113,Histogenetics, Ossining, USA
C*07:616HG00013718MG603114,Histogenetics, Ossining, USA
C*07:617HG00013891MG701528,Histogenetics, Ossining, USA
C*07:618HG00014123,HG00014166,HG00014173,HG00014124MG701535,MG724865,MG724872,MG701536,Histogenetics, Ossining, USA
C*07:619HG00014126,HG00014177,HG00013900MG701538,MG724876,MG724856,Histogenetics, Ossining, USA
C*07:620HG00014153MG701565,Histogenetics, Ossining, USA
C*07:621HG00014179,HG00014187,HG00014199MG724878,MG724886,MG724898,Histogenetics, Ossining, USA
C*07:622HG00014185MG724884,Histogenetics, Ossining, USA
C*07:623HG00014193,HG00014481,HG00014482,HG00013698,HG00013699MG724892,MG756855,MG756856,MG584046,MG584047,Histogenetics, Ossining, USA
C*07:624HG00014457MG756829,Histogenetics, Ossining, USA
C*07:625HG00014473MG756847,Histogenetics, Ossining, USA
C*07:626DKMS-LSL-C-2387LT989975,DKMS Life Sciences Lab, Dresden, Germany
C*07:627DKMS-LSL-C-2437LT990025,DKMS Life Sciences Lab, Dresden, Germany
C*08:02:01:03DKMS-LSL-C-2393LT989981,DKMS Life Sciences Lab, Dresden, Germany
C*08:02:16HG00013720MG603116,Histogenetics, Ossining, USA
C*08:02:17HG00014188MG724887,Histogenetics, Ossining, USA
C*08:62:02HG00013715MG584063,Histogenetics, Ossining, USA
C*08:144:02HG00013719,HG00013736MG603115,MG603132,Histogenetics, Ossining, USA
C*08:162HG00013708MG584056,Histogenetics, Ossining, USA
C*08:163HG00013724MG603120,Histogenetics, Ossining, USA
C*08:164HG00014475,HG00014135MG756849,MG701547,Histogenetics, Ossining, USA
C*08:165JRC-D44-116LC371760,Masahiro Satake, Tokyo, Japan
C*08:166DKMS-LSL-C-2407,DKMS-LSL-C-2406LT989995,LT989994,DKMS Life Sciences Lab, Dresden, Germany
C*12:02:16HG00013721MG603117,Histogenetics, Ossining, USA
C*12:02:17JRC-D01-490LC371762,Masahiro Satake, Tokyo, Japan
C*12:03:44HG00014494MG756869,Histogenetics, Ossining, USA
C*12:03:45DKMS-LSL-C-2416LT990004,DKMS Life Sciences Lab, Dresden, Germany
C*12:03:46DKMS-LSL-C-2402LT989990,DKMS Life Sciences Lab, Dresden, Germany
C*12:03:47DKMS-LSL-C-2432LT990020,DKMS Life Sciences Lab, Dresden, Germany
C*12:223HG00013701MG584049,Histogenetics, Ossining, USA
C*12:224HG00013704MG584052,Histogenetics, Ossining, USA
C*12:225HG00013705MG584053,Histogenetics, Ossining, USA
C*12:226HG00013723MG603119,Histogenetics, Ossining, USA
C*12:227HG00013742,HG00013740MG603138,MG603136,Histogenetics, Ossining, USA
C*12:228HG00013890MG701525,Histogenetics, Ossining, USA
C*12:229HG00014493MG756868,Histogenetics, Ossining, USA
C*12:230DKMS-LSL-C-2431LT990019,DKMS Life Sciences Lab, Dresden, Germany
C*12:231DKMS-LSL-C-2449LT990037,DKMS Life Sciences Lab, Dresden, Germany
C*12:232NDKMS-LSL-C-2447LT990035,DKMS Life Sciences Lab, Dresden, Germany
C*12:233DKMS-LSL-C-2435LT990023,DKMS Life Sciences Lab, Dresden, Germany
C*14:02:01:06DKMS-LSL-C-2409LT989997,DKMS Life Sciences Lab, Dresden, Germany
C*14:02:21HG00014149MG701561,Histogenetics, Ossining, USA
C*14:94HG00014156MG701568,Histogenetics, Ossining, USA
C*14:95HG00014496,HG00014497,HG00014495MG756871,MG756872,MG756870,Histogenetics, Ossining, USA
C*14:96DKMS-LSL-C-2392LT989980,DKMS Life Sciences Lab, Dresden, Germany
C*15:02:32JXZJ20171101004MH054903,Faming Zhu, Hangzhou, China
C*15:23:02HG00013732MG603128,Histogenetics, Ossining, USA
C*15:152:02HG00013703MG584051,Histogenetics, Ossining, USA
C*15:153HG00013714MG584062,Histogenetics, Ossining, USA
C*15:154HG00013904MG724860,Histogenetics, Ossining, USA
C*15:155DKMS-LSL-C-2414LT990002,DKMS Life Sciences Lab, Dresden, Germany
C*16:01:23DKMS-LSL-C-2436LT990024,DKMS Life Sciences Lab, Dresden, Germany
C*16:02:15HG00014155MG701567,Histogenetics, Ossining, USA
C*16:118HG00013893MG701530,Histogenetics, Ossining, USA
C*16:119BST1339255MG999586,Francesc Rudilla, Barcelona, Spain
C*16:120DKMS-LSL-C-2399LT989987,DKMS Life Sciences Lab, Dresden, Germany
C*16:121DKMS-LSL-C-2438LT990026,DKMS Life Sciences Lab, Dresden, Germany
C*16:122DKMS-LSL-C-2419LT990007,DKMS Life Sciences Lab, Dresden, Germany
C*17:40DKMS-LSL-C-2417LT990005,DKMS Life Sciences Lab, Dresden, Germany
DPB1*02:01:22DKMS-LSL-DPB1-1709,DKMS-LSL-DPB1-1708LT882733,LT882732,DKMS Life Sciences Lab, Dresden, Germany
DPB1*04:01:37DKMS-LSL-DPB1-1723LT882747,DKMS Life Sciences Lab, Dresden, Germany
DPB1*699:01DKMS-LSL-DPB1-1760,DKMS-LSL-DPB1-1759LT883486,LT883485,DKMS Life Sciences Lab, Dresden, Germany
DPB1*700:01NDKMS-LSL-DPB1-1711,DKMS-LSL-DPB1-1710LT882735,LT882734,DKMS Life Sciences Lab, Dresden, Germany
DPB1*701:012018-1186MG972812,Mauro Castagnetta, Genova, Italy
DPB1*702:0171708570MG954069,Stephane Buhler, Geneva, Switzerland
DQB1*02:10671703751MG954063,Stephane Buhler, Geneva, Switzerland
DQB1*03:280H180019MG993320,Faming Zhu, Hangzhou, China
DQB1*05:16093030789MH051245,Daniel Fuerst, Ulm, Germany
G*01:22TXR23MG831330,Vanessa Hauer, Curitiba, Brazil
MICB*002:03WN127MG913793,Tian Wei, Urumqi, China

Table 2: Confirmatory Sequences


Sequence Cell identification Accession number Submitting author
A*01:01:83HG00013861,HG00013959,HG00013982,HG00014022,HG00014052,MG724752,MG701434,MG701457,MG724778,MG724808,Histogenetics, Ossining, USA
A*01:04NMYE 2001,LT976483,DKMS Life Sciences Lab, Dresden, Germany
A*01:09:01:02NT02127,NT02128,MH036498,Carolyn K Hurley, Washington DC, USA
A*01:69:01NT02131,MH036496,Carolyn K Hurley, Washington DC, USA
A*02:16TUBO,LT976476,DKMS Life Sciences Lab, Dresden, Germany
A*02:6748973_maas04,LT985838,Marcel GJ Tilanus, Maastricht, The Netherlands
A*02:686HG00013630,HG00013855,HG00013859,HG00013955,HG00013972,HG00013975,HG00013997,HG00014013,HG00014015,HG00014026,HG00014028,HG00014032,HG00014039,HG00014047,HG00014050,MG603070,MG701417,MG724750,MG701430,MG701447,MG701450,MG701472,MG724769,MG724771,MG724782,MG724784,MG724788,MG724795,MG724803,MG724806,Histogenetics, Ossining, USA
A*02:686HG00013862,HG00014063,MG724753,MG724819,Histogenetics, Ossining, USA
A*02:686HG00015471,HG00015496,HG00015497,MG930525,MG930550,MG930551,Histogenetics, Ossining, USA
A*02:70471708599,MG954070,Stephane Buhler, Geneva, Switzerland
A*02:70717228634,MH051244,Daniel Fuerst, Ulm, Germany
A*03:01:034993,LT976502,Marcel GJ Tilanus, Maastricht, The Netherlands
A*03:22:01NT02132,MH036497,Carolyn K Hurley, Washington DC, USA
A*03:25NT02133,MH036494,Carolyn K Hurley, Washington DC, USA
A*03:88NT02130,MH036495,Carolyn K Hurley, Washington DC, USA
A*03:103:02HG00013624,MG584015,Histogenetics, Ossining, USA
A*03:288HG00013993,HG00013994,MG701468,MG701469,Histogenetics, Ossining, USA
A*11:172Q2,MG782888,Yunping Xu, Shenzhen, China
A*11:263HG00014058,HG00014061,MG724814,MG724817,Histogenetics, Ossining, USA
A*24:02:2648986_maas18,LT985839,Marcel GJ Tilanus, Maastricht, The Netherlands
A*25:04NT02129,MH036499,Carolyn K Hurley, Washington DC, USA
A*26:03:01S.M.,LT976481,DKMS Life Sciences Lab, Dresden, Germany
A*30:03JS,LT976478,DKMS Life Sciences Lab, Dresden, Germany
A*32:103HG00013949,HG00013950,HG00014048,HG00014057,MG701424,MG701425,MG724804,MG724813,Histogenetics, Ossining, USA
A*33:134HG00013999,MG701474,Histogenetics, Ossining, USA
A*66:01:01:01IDF,LT976477,DKMS Life Sciences Lab, Dresden, Germany
A*68:11NMYE 2003,LT976484,DKMS Life Sciences Lab, Dresden, Germany
A*74:25NT02134 ,MH036500,Carolyn K Hurley, Washington DC, USA
B*07:02:53BST13256570,MH045582,Francesc Rudilla, Barcelona, Spain
B*07:09NT02087,MG986062,Carolyn K Hurley, Washington DC, USA
B*07:2048988_maas20,LT985837,Marcel GJ Tilanus, Maastricht, The Netherlands
B*08:12:01NT02098,MH015216,Carolyn K Hurley, Washington DC, USA
B*08:188HG00014097,MG701517,Histogenetics, Ossining, USA
B*14:3148992_maas24,LT985858,Marcel GJ Tilanus, Maastricht, The Netherlands
B*15:31NT02088,MG986063,Carolyn K Hurley, Washington DC, USA
B*15:33NT02089,MG986064,Carolyn K Hurley, Washington DC, USA
B*15:110NT02097,MH015214,Carolyn K Hurley, Washington DC, USA
B*15:178S10,MG782891,Yunping Xu, Shenzhen, China
B*18:7348996_maas28,LT985841,Marcel GJ Tilanus, Maastricht, The Netherlands
B*35:90NT02091,MG986066,Carolyn K Hurley, Washington DC, USA
B*35:23148998_maas30,LT985852,Marcel GJ Tilanus, Maastricht, The Netherlands
B*37:70HG00013683,MG603101,Histogenetics, Ossining, USA
B*39:131HG00014100,MG701520,Histogenetics, Ossining, USA
B*40:16NT02092,MG986067,Carolyn K Hurley, Washington DC, USA
B*40:42NT02093,NT02094,MG986068,Carolyn K Hurley, Washington DC, USA
B*40:54Q1,MG782889,Yunping Xu, Shenzhen, China
B*41:5516220512,MG973020,Daniel Fuerst, Ulm, Germany
B*47:03NT02095,MG986069,Carolyn K Hurley, Washington DC, USA
B*48:03:02NT02105,MH015215,Carolyn K Hurley, Washington DC, USA
B*48:06NT02096,MG986070,Carolyn K Hurley, Washington DC, USA
B*50:31Q5,MG782890,Yunping Xu, Shenzhen, China
B*50:55HG00014092,MG701512,Histogenetics, Ossining, USA
B*56:55:01:0271708646,MG954071,Stephane Buhler, Geneva, Switzerland
C*01:126DKMS-LSL-C-2430,LT990018,DKMS Life Sciences Lab, Dresden, Germany
C*02:02:02:03DKMS-LSL-C-2388,LT989976,DKMS Life Sciences Lab, Dresden, Germany
C*02:10:032017-2931,MG972811,Mauro Castagnetta, Genova, Italy
C*02:60DKMS-LSL-C-2440,LT990028,DKMS Life Sciences Lab, Dresden, Germany
C*02:98DKMS-LSL-C-2404,LT989992,DKMS Life Sciences Lab, Dresden, Germany
C*03:02:1717ZZ0352,LC371654,LC371402,Li-Qun Zhang, Beijing, China
C*03:4449002_maas35,LT989888,Marcel GJ Tilanus, Maastricht, The Netherlands
C*03:359HG00014178,HG00014186,HG00014169,HG00014182,HG00014200,MG724877,MG724885,MG724868,MG724881,MG724899,Histogenetics, Ossining, USA
C*03:362HG00014485,HG00014486,MG756859,MG756860,Histogenetics, Ossining, USA
C*05:1349003_maas36,LT985859,Marcel GJ Tilanus, Maastricht, The Netherlands
C*05:154NHG00013716,MG584064,Histogenetics, Ossining, USA
C*05:158BST008,MG967419,Francesc Rudilla, Barcelona, Spain
C*07:4649005_maas38,LT985840,Marcel GJ Tilanus, Maastricht, The Netherlands
C*07:250NT02086,MG980571,Carolyn K Hurley, Washington DC, USA
C*07:526DKMS-LSL-C-2412,LT990000,DKMS Life Sciences Lab, Dresden, Germany
C*07:595HG00014165,HG00014189,MG724864,MG724888,Histogenetics, Ossining, USA
C*07:610HG00013903,MG724859,Histogenetics, Ossining, USA
C*07:612HG00013906,MG724862,Histogenetics, Ossining, USA
C*07:613DKMS-LSL-C-2441,LT990029,DKMS Life Sciences Lab, Dresden, Germany
C*08:148HG00014190,MG724889,Histogenetics, Ossining, USA
C*08:152HG00013739,MG603135,Histogenetics, Ossining, USA
C*14:02:19DKMS-LSL-C-2429,LT990017,DKMS Life Sciences Lab, Dresden, Germany
C*15:02:31HG00013898,MG724853,Histogenetics, Ossining, USA
C*15:1149007_maas40,LT985876,Marcel GJ Tilanus, Maastricht, The Netherlands
C*15:13:01:02HG00014125,MG701537,Histogenetics, Ossining, USA
C*17:38HG00013895,MG701533,Histogenetics, Ossining, USA
C*17:38HG00014499,MG756875,Histogenetics, Ossining, USA
DPB1*416:01:01:02DKMS-LSL-DPB1-1775,LT883501,DKMS Life Sciences Lab, Dresden, Germany
DPB1*652:0171705809,MG954067,Stephane Buhler, Geneva, Switzerland
DQB1*03:01:03NT02135,MH042876,Carolyn K Hurley, Washington DC, USA
DQB1*03:02:02NT02137,NT02136,MH042877,Carolyn K Hurley, Washington DC, USA
DQB1*03:03:11NT02138,MH042878,Carolyn K Hurley, Washington DC, USA
DQB1*03:08NT02139,MH042879,Carolyn K Hurley, Washington DC, USA
DQB1*03:72NT02140,NT02141,MH042880,Carolyn K Hurley, Washington DC, USA
DQB1*06:03:01:02#15310-LN,LT976480,DKMS Life Sciences Lab, Dresden, Germany
DQB1*06:07:01NT02099,MH017034,Carolyn K Hurley, Washington DC, USA
DQB1*06:08:01NT02100,NT02101,MH017035,Carolyn K Hurley, Washington DC, USA
DQB1*06:09:02NT02102,MH017036,Carolyn K Hurley, Washington DC, USA
DQB1*06:10NT02104,NT02103,MH017037,Carolyn K Hurley, Washington DC, USA
DQB1*06:11:01NT02106,NT02107,MH017038,Carolyn K Hurley, Washington DC, USA
DQB1*06:12NT02108,MH017039,Carolyn K Hurley, Washington DC, USA
DQB1*06:13:01NT02109,MH017049,Carolyn K Hurley, Washington DC, USA
DQB1*06:14:01NT02111,NT02110,MH017040,Carolyn K Hurley, Washington DC, USA
DQB1*06:14:02NT02112,MH017041,Carolyn K Hurley, Washington DC, USA
DQB1*06:19:02NT02114,MH017043,Carolyn K Hurley, Washington DC, USA
DQB1*06:20NT02115,MH017044,Carolyn K Hurley, Washington DC, USA
DQB1*06:33NT02116,NT02117,MH017045,Carolyn K Hurley, Washington DC, USA
DQB1*06:46NT02119,NT02118,MH017046,Carolyn K Hurley, Washington DC, USA
DQB1*06:48NT02120,MH017047,Carolyn K Hurley, Washington DC, USA
DQB1*06:49NT02121,NT02122,MH017048,Carolyn K Hurley, Washington DC, USA
DQB1*06:110MGM2_118357,MG672522,Vibhuti Rambani, Bangalore, India

Table 3: Recently Published Sequences


Sequence References
A*24:391(3)
A*26:118(4)
A*33:138(5)
B*07:305(6)
B*14:58(7)
B*18:37:02(8)
B*40:323(9)
B*40:333(10)
B*40:338N(11)
C*01:136(12)
C*03:03:35(13)
C*15:151(14)
C*16:116(15)
DPB1*647:01(16)
DRB1*07:83(17)
DRB1*14:177(9)
DQB1*02:65(4)
DQB1*03:200(9)
DQB1*04:01:05(18)
DQB1*06:205(9)
DPB1*05:01:07(4)

References:

  1. Marsh SGE, Albert ED, Bodmer WF, et al. Nomenclature for factors of the HLA system, 2010. Tissue Antigens 2010 75: 291-455.
  2. Robinson J, Halliwell JA, Hayhurst JD, Flicek P, Parham P, Marsh SGE. The IPD and IMGT/HLA database: allele variant databases. Nucleic Acids Res 2015: 43: 423-31.
  3. Guidicelli G, Elsermans V, Top I, et al.Characterization of the novel HLA-A*24:391 allele by sequencing-based typing. HLA 2018 91: 292-293.
  4. Park BG, Kim SK, Im J, et al.Three novel HLA alleles in discovered in Koreans, HLA-A*26:118, -DQB1*02:65 and -DPB1*05:01:07 HLA 2018 91: 293-294.
  5. Huo MR, Yu Y, Xi B, et al.Identification of a novel HLA-A*33 allele, HLA-A*33:138. HLA 2018 91: 294-295.
  6. Guidicelli G, Elsermans V, Top I, et al.Characterization of the novel HLA-B*07:305 allele by sequencing-based typing. HLA 2018 91: 296-297.
  7. Ingrassia F, Mistretta S, Bavetta R, et al.Characterization of the novel allele: B*14:58. HLA 2018 91: 297-298.
  8. Burek Kamernaric M, Maskalan M, Drabbels J, et al.Identification of the novel HLA-B*18:37:02 allele in a Croatian individual. HLA 2018 91: 299-300.
  9. Park BG, Kim SK, Im J, et al.Four novel alleles in Korean individuals, HLA-B*40:323, -DRB1*14:177, -DQB1*03:200 and -DQB1*06:205 HLA 2018 91: 300-301.
  10. Zhao SX, Han ZD, He J, et al.Identification of the novel HLA-B*40:333 allele by polymerase chain reaction sequence-based typing. HLA 2018 91: 302-303.
  11. He J, Dong LN, Wang F, et al.The novel null allele, HLA-B*40:338N, was identified in a Chinese leukemia patient. HLA 2018 91: 303-305.
  12. Zingsem J, Hauck-Dlimi B, Dullinger K, et al.Identification of the novel allele, HLA-C*01:136, in a German cord blood donor originating from Azerbaijan. HLA 2018 91: 305-306.
  13. Park BG, Oh HB, Im J, et al.Identification of the novel HLA-C*03 allele, HLA-C*03:03:35. HLA 2018 91: 306-307.
  14. Yang KL, Lin PY.Identification of a novel HLA-C allele, HLA-C*15:151, in a Taiwanese individual. HLA 2018 91: 308-309.
  15. Guidicelli G, Elsermans V, Top I, et al.Characterization of the novel HLA-C*16:116 allele by sequencing-based typing. HLA 2018 91: 309-311.
  16. Fornaciari S, Sciarrino R, Biagini C, et al.Identification of the new HLA-DPB1 allele, DPB1*647:01, in an Italian patient with leukemia. HLA 2018 91: 311-312.
  17. Balas A, Alenda R, Moreno-Hidalgo MA, et al.Characterization of a novel HLA-DRB1*07 allele, DRB1*07:83. HLA 2018 91: 313-314.
  18. Park BG, Oh HB, Im J, et al.A new HLA-DQB1*04 allele, HLA-DQB1*04:01:05, identified in a Korean individual. HLA 2018 91: 312-313.