Nomenclature for Factors of the HLA System

August 2016

Compiled by Steven G. E. Marsh for the WHO Nomenclature Committee for Factors of the HLA System.

Published In:

The following sequences have been submitted to the Nomenclature Committee since the July 2016 nomenclature update and, following agreed policy, have been assigned official allele designations. Full details of all sequences will be published in a forthcoming report.

Below are listed the newly assigned sequences (Table 1) and confirmations of previously reported sequences (Table 2). The accession number of each sequence is given and these can be used to retrieve the sequence files from the EMBL, GenBank or DDBJ data libraries. Although accession numbers have been assigned by the data-libraries and most sequences are already available, there is still the possibility that an author may not yet have allowed the sequence to be released; in such a case you will have to contact the submitting author directly. Additional information pertaining to new sequences is often included in the publications describing these alleles; a listing of recent publications that describe new HLA sequences is given in Table 3. The sequence of C*17:01:01:01 has been found to be in error and the corrected sequence is identical to C*17:01:01:02, C*17:01:01:01 has therefore been abandoned.

All new and confirmatory sequences should now be submitted directly to the WHO Nomenclature Committee for Factors of the HLA System via the IMGT/HLA Database using the sequence submission tool provided (2). The IMGT/HLA Database may be accessed via the world wide web at : http://ww.ebi.ac.uk/imgt/hla/.

Table 1: New Sequences


SequenceCell identificationAccession numberSubmitting author
A*01:2121450-3131-6 KX622665,Yuxin Yin, Los Angeles, USA
A*01:213M-374849LT605167,Kristin Kipper, Martinsried, Germany
A*02:01:01:09D041423LN901401,Edward KL Yang, Hualien, Taiwan
A*02:591:02M-637541LT605169,Kristin Kipper, Martinsried, Germany
A*02:641HG00012236KX545278,Histogenetics, Ossining, USA
A*02:6421429-2056-0 ,1429-2486-9 ,1479-1178-8 KX622669,KX622670,KX622671,Yuxin Yin, Los Angeles, USA
A*02:643NJF25732KX245151,Raquel Fabreti, Belo Horizonte, Brazil
A*11:2471454-5757-8 KX622667,Yuxin Yin, Los Angeles, USA
A*11:2481729-1880-7 KX622668,Yuxin Yin, Los Angeles, USA
A*11:249DaBC_04KX278826,Yu Weijian, DaLian, China
A*24:02:981724-3465-6 ,1429-2294-7 ,1465-2630-6 ,1732-7757-5 ,1737-7763-2 KX622659,KX622660,KX622661,KX622662,KX622663,Yuxin Yin, Los Angeles, USA
A*24:3521744-1843-4 KX622672,Yuxin Yin, Los Angeles, USA
A*24:3531459-9658-3 KX622675,Yuxin Yin, Los Angeles, USA
A*24:354DaBC_05KX278827,Yu Weijian, DaLian, China
A*26:01:401673-3677-5 KX622673,Yuxin Yin, Los Angeles, USA
A*26:128M-474143LT596578,Kristin Kipper, Martinsried, Germany
A*30:104HG00012235KX545279,Histogenetics, Ossining, USA
A*30:1051674-4496-7 KX622676,Yuxin Yin, Los Angeles, USA
A*32:91M-425534LT605170,Kristin Kipper, Martinsried, Germany
A*33:01:091729-9501-1 KX622664,Yuxin Yin, Los Angeles, USA
A*33:117212032KX611133,Raquel Fabreti, Belo Horizonte, Brazil
A*34:17HG00012233KX578045,Histogenetics, Ossining, USA
A*68:151HG00012234KX578046,Histogenetics, Ossining, USA
A*68:1521268-2045-5 KX622674,Yuxin Yin, Los Angeles, USA
B*07:278HG00012089,HG00012090,HG00012091KX173363,KX173364,KX173365,Histogenetics, Ossining, USA
B*07:279HG00012092KX173349,Histogenetics, Ossining, USA
B*07:280HG00012099KX173359,Histogenetics, Ossining, USA
B*07:281HG00012247KX499421,Histogenetics, Ossining, USA
B*07:2821464-9206-1 KX622688,Yuxin Yin, Los Angeles, USA
B*07:283M-468075LT605171,Kristin Kipper, Martinsried, Germany
B*14:55HG00012095KX425968,Histogenetics, Ossining, USA
B*15:27:04HG00012245KX545281,Histogenetics, Ossining, USA
B*15:363:02HG00012083KX266771,Histogenetics, Ossining, USA
B*15:402HG00012084KX266768,Histogenetics, Ossining, USA
B*15:403HG00012085KX173358,Histogenetics, Ossining, USA
B*18:130M-406030LT605172,Kristin Kipper, Martinsried, Germany
B*27:155HG00012240KX499425,Histogenetics, Ossining, USA
B*27:156015874907LT595713,Dominique Masson, La Tronche, France
B*38:64HG00012096,HG00012097,HG00012098KX425964,KU350645,KX425962,Histogenetics, Ossining, USA
B*39:120HG00012094KX173360,Histogenetics, Ossining, USA
B*40:01:48RDP-CH1-20KX686514,Carolyn K Hurley, Washington DC, USA
B*40:342JRC-D79-068LC171642,Masahiro Satake, Tokyo, Japan
B*40:343143964KX611135,Raquel Fabreti, Belo Horizonte, Brazil
B*40:344DaBC_03KX278825,Yu Weijian, DaLian, China
B*41:48102748KX611134,Raquel Fabreti, Belo Horizonte, Brazil
B*41:49M-638693LT605168,Kristin Kipper, Martinsried, Germany
B*42:24265986KX611138,Raquel Fabreti, Belo Horizonte, Brazil
B*44:2551739-6033-7 KX622686,Yuxin Yin, Los Angeles, USA
B*46:69HG00012082KX173351,Histogenetics, Ossining, USA
B*48:41HG00012093KU350644,Histogenetics, Ossining, USA
B*49:45M-393372LT605173,Kristin Kipper, Martinsried, Germany
B*50:48260908KX245150,Raquel Fabreti, Belo Horizonte, Brazil
B*51:63:02HG00012088KX425959,Histogenetics, Ossining, USA
B*51:210HG00012246KX578058,Histogenetics, Ossining, USA
B*54:36HG00012086KX425963,Histogenetics, Ossining, USA
B*55:791751-7333-5 KX622687,Yuxin Yin, Los Angeles, USA
B*57:85M-441789LT605174,Kristin Kipper, Martinsried, Germany
B*58:83HG00012244KX499419,Histogenetics, Ossining, USA
C*01:126HG00012103KX425972,Histogenetics, Ossining, USA
C*01:1271710-5087-5 KX622683,Yuxin Yin, Los Angeles, USA
C*02:121NHG00012252KX455873,Histogenetics, Ossining, USA
C*03:04:521720-7527-7KX622677,Yuxin Yin, Los Angeles, USA
C*03:04:53RDP-CH4-82KX686518,Carolyn K Hurley, Washington DC, USA
C*03:328HG00012137KX425985,Histogenetics, Ossining, USA
C*03:329HG00012138KX425984,Histogenetics, Ossining, USA
C*03:330HG00012140KX173379,Histogenetics, Ossining, USA
C*03:331HG00012141KX173376,Histogenetics, Ossining, USA
C*03:332HG00012142KX289898,Histogenetics, Ossining, USA
C*03:333HG00012143KX173384,Histogenetics, Ossining, USA
C*03:334HG00012144KX425975,Histogenetics, Ossining, USA
C*03:3351469-9772-1 KX622681,Yuxin Yin, Los Angeles, USA
C*03:336M-630315LT605175,Kristin Kipper, Martinsried, Germany
C*04:01:771461-8629-1 KX622679,Yuxin Yin, Los Angeles, USA
C*04:01:781729-0597-8 KX622682,Yuxin Yin, Los Angeles, USA
C*04:01:79RDP-CH4-08KX686516,Carolyn K Hurley, Washington DC, USA
C*04:223:02HG00012149KX266774,Histogenetics, Ossining, USA
C*04:249HG00012145KX289902,Histogenetics, Ossining, USA
C*04:250HG00012146,HG00012147KX173386,KX266783,Histogenetics, Ossining, USA
C*04:251HG00012148KX266777,Histogenetics, Ossining, USA
C*04:252HG00012150KX289901,Histogenetics, Ossining, USA
C*04:253NHG00012151KX229697,Histogenetics, Ossining, USA
C*04:254HG00012152,HG00012153KX173373,KX425976,Histogenetics, Ossining, USA
C*04:255NM-388611LT605177,Kristin Kipper, Martinsried, Germany
C*05:136HG00012117KU360752,Histogenetics, Ossining, USA
C*05:13716536532KX458015,Antonio Balas, Madrid, Spain
C*05:138HG00012251KX545288,Histogenetics, Ossining, USA
C*06:191HG00012105KX289904,Histogenetics, Ossining, USA
C*06:192HG00012106KX289896,Histogenetics, Ossining, USA
C*07:534HG00012107KX425974,Histogenetics, Ossining, USA
C*07:535HG00012108KX289897,Histogenetics, Ossining, USA
C*07:536HG00012109KX173385,Histogenetics, Ossining, USA
C*07:537HG00012110KX173382,Histogenetics, Ossining, USA
C*07:538HG00012112KX425987,Histogenetics, Ossining, USA
C*07:539HG00012114KX266784,Histogenetics, Ossining, USA
C*07:540HG00012115KU350647,Histogenetics, Ossining, USA
C*07:541HG00012248KX499426,Histogenetics, Ossining, USA
C*07:542HG00012249KX499427,Histogenetics, Ossining, USA
C*07:5431713-8278-1 KX622678,Yuxin Yin, Los Angeles, USA
C*07:544D225190LT605075,Edward KL Yang, Hualien, Taiwan
C*07:545M-568695LT605176,Kristin Kipper, Martinsried, Germany
C*08:137HG00012119KX173374,Histogenetics, Ossining, USA
C*12:189HG00012120KX266778,Histogenetics, Ossining, USA
C*12:190HG00012124KX266776,Histogenetics, Ossining, USA
C*12:191HG00012125KX266775,Histogenetics, Ossining, USA
C*12:192HG00012250KX578050,Histogenetics, Ossining, USA
C*12:193M-616269LT605178,Kristin Kipper, Martinsried, Germany
C*15:130DKMS-LSL-C-667LN912934,DKMS Life Sciences Lab, Dresden, Germany
C*15:131HG00012134KX173378,Histogenetics, Ossining, USA
C*16:99HG00012131KX289899,Histogenetics, Ossining, USA
C*16:1001732-8556-0 KX622680,Yuxin Yin, Los Angeles, USA
C*17:34HG00012129KX173375,Histogenetics, Ossining, USA
DPA1*02:07IND-1KU239966,Konstantinos Barsakis, Palo Alto, USA
DPB1*10:01:03HG00012227KX173408,Histogenetics, Ossining, USA
DPB1*65:01:02HG00012215KX426015,Histogenetics, Ossining, USA
DPB1*90:01:02HG00012228,HG00012229KX173413,KX173414,Histogenetics, Ossining, USA
DPB1*586:01HG00012209KX289918,Histogenetics, Ossining, USA
DPB1*587:01HG00012210KX426020,Histogenetics, Ossining, USA
DPB1*588:01HG00012211KU360780,Histogenetics, Ossining, USA
DPB1*589:01HG00012212KX266805,Histogenetics, Ossining, USA
DPB1*590:01HG00012213KX266809,Histogenetics, Ossining, USA
DPB1*591:01HG00012216KX266806,Histogenetics, Ossining, USA
DPB1*592:01HG00012217KX173417,Histogenetics, Ossining, USA
DPB1*593:01HG00012218KX173420,Histogenetics, Ossining, USA
DPB1*594:01HG00012219KX426016,Histogenetics, Ossining, USA
DPB1*595:01HG00012221KX266804,Histogenetics, Ossining, USA
DPB1*596:01HG00012224KX289921,Histogenetics, Ossining, USA
DPB1*597:01HG00012231KX426017,Histogenetics, Ossining, USA
DPB1*598:01NHG00012261KX455875,Histogenetics, Ossining, USA
DPB1*599:01HG00012262KX499435,Histogenetics, Ossining, USA
DPB1*600:01HG00012263KX545293,Histogenetics, Ossining, USA
DPB1*601:01HG00012264KX578064,Histogenetics, Ossining, USA
DQA1*01:14RDP-CH4-28KX686517,Carolyn K Hurley, Washington DC, USA
DQA1*05:05:01:07BEL7MONKU319359,Konstantinos Barsakis, Palo Alto, USA
DQA1*05:05:01:08C047KU319360,Konstantinos Barsakis, Palo Alto, USA
DQB1*02:02:031732-9695-5 KX622650,Yuxin Yin, Los Angeles, USA
DQB1*03:02:20JF34865KX646399,Raquel Fabreti, Belo Horizonte, Brazil
DQB1*03:235HG00012155KU360761,Histogenetics, Ossining, USA
DQB1*03:236HG00012156KU360759,Histogenetics, Ossining, USA
DQB1*03:237NHG00012163KU360756,Histogenetics, Ossining, USA
DQB1*03:238HG00012164KX289907,Histogenetics, Ossining, USA
DQB1*03:239HG00012165KU350651,Histogenetics, Ossining, USA
DQB1*03:240HG00012166,HG00012167,HG00012168,HG00012169,HG00012170KX289910,KX289911,KX289912,KX289913,KX425996,Histogenetics, Ossining, USA
DQB1*03:2411728-4535-6 KX622647,Yuxin Yin, Los Angeles, USA
DQB1*03:2421723-5417-7 KX622648,Yuxin Yin, Los Angeles, USA
DQB1*03:243244744,243991KX430222,Raquel Fabreti, Belo Horizonte, Brazil
DQB1*04:38HG00012177KX425997,Histogenetics, Ossining, USA
DQB1*05:125HG00012161KX173390,Histogenetics, Ossining, USA
DQB1*05:126HG00012171KX266785,Histogenetics, Ossining, USA
DQB1*05:127HG00012172KX425990,Histogenetics, Ossining, USA
DQB1*05:128NHG00012173KX229700,Histogenetics, Ossining, USA
DQB1*05:129HG00012175KX266786,Histogenetics, Ossining, USA
DQB1*05:130HG00012254KX578060,Histogenetics, Ossining, USA
DQB1*06:212HG00012159,HG00012160KX425994,KX425995,Histogenetics, Ossining, USA
DQB1*06:213HG00012178KX425991,Histogenetics, Ossining, USA
DQB1*06:214HG00012253KX499428,Histogenetics, Ossining, USA
DRB1*01:76HG00012196KX173399,Histogenetics, Ossining, USA
DRB1*01:77M-507122LT605179,Kristin Kipper, Martinsried, Germany
DRB1*03:134HG00012186KX173400,Histogenetics, Ossining, USA
DRB1*03:135HG00012195KX426004,Histogenetics, Ossining, USA
DRB1*03:136HG00012256KX499429,Histogenetics, Ossining, USA
DRB1*03:1371730-9667-8 KX622653,Yuxin Yin, Los Angeles, USA
DRB1*04:216HG00012183KX173397,Histogenetics, Ossining, USA
DRB1*04:217HG00012184KX173396,Histogenetics, Ossining, USA
DRB1*07:01:20015765318LT595712,Dominique Masson, La Tronche, France
DRB1*07:01:211734-1875-7 KX622652,Yuxin Yin, Los Angeles, USA
DRB1*07:73HG00012182KX173401,Histogenetics, Ossining, USA
DRB1*07:74HG00012255KX545284,Histogenetics, Ossining, USA
DRB1*10:18HG00012185KX426003,Histogenetics, Ossining, USA
DRB1*10:191741-8896-1 KX622656,Yuxin Yin, Los Angeles, USA
DRB1*11:04:14259284KX646401,Raquel Fabreti, Belo Horizonte, Brazil
DRB1*11:203HG00012192KX266795,Histogenetics, Ossining, USA
DRB1*11:204HG00012193KU360764,Histogenetics, Ossining, USA
DRB1*11:205HG00012194KX289915,Histogenetics, Ossining, USA
DRB1*13:02:131741-8570-2 KX622655,Yuxin Yin, Los Angeles, USA
DRB1*13:221HG00012188KX289914,Histogenetics, Ossining, USA
DRB1*13:222HG00012189KX425998,Histogenetics, Ossining, USA
DRB1*13:223HG00012190KU360765,Histogenetics, Ossining, USA
DRB1*13:224HG00012191KU360766,Histogenetics, Ossining, USA
DRB1*13:225HG00012258KX499430,Histogenetics, Ossining, USA
DRB1*14:02:03HG00012187KX289916,Histogenetics, Ossining, USA
DRB1*15:01:31HG00012180KX266792,Histogenetics, Ossining, USA
DRB1*15:01:321712-6075-5 KX622657,Yuxin Yin, Los Angeles, USA
DRB1*16:43HG00012181KX426002,Histogenetics, Ossining, USA
DRB1*16:44015809650LT595711,Dominique Masson, La Tronche, France
DRB3*01:40NHG00012259KX455874,Histogenetics, Ossining, USA
DRB3*02:55NHG00012197KX229698,Histogenetics, Ossining, USA
DRB3*02:56HG00012198KX289917,Histogenetics, Ossining, USA
DRB3*02:57HG00012200KU360767,Histogenetics, Ossining, USA
DRB4*01:42HG00012202KX426012,Histogenetics, Ossining, USA
DRB4*01:43HG00012203KU360775,Histogenetics, Ossining, USA
DRB4*01:44HG00012204,HG00012205KX266801,KX426011,Histogenetics, Ossining, USA
DRB5*02:15HG00012207KU360778,Histogenetics, Ossining, USA
E*01:01:01:06SRSA5073-EP44KX709623,Yunping Xu, Shenzhen, China
E*01:01:01:07SRSA5097-EP71KX709625,Yunping Xu, Shenzhen, China

Table 2: Confirmatory Sequences


Sequence Cell identification Accession number Submitting author
A*02:29NT01419,KX583636,Carolyn K Hurley, Washington DC, USA
A*02:58NT01420,KX583635,Carolyn K Hurley, Washington DC, USA
A*03:01:02RDP-CH1-53,KX686515,Carolyn K Hurley, Washington DC, USA
A*03:01:03NT01421,KX583634,Carolyn K Hurley, Washington DC, USA
A*03:57NT01417,KX583637,Carolyn K Hurley, Washington DC, USA
A*23:05NT01422,KX583633,Carolyn K Hurley, Washington DC, USA
A*24:23NT01423,KX583632,Carolyn K Hurley, Washington DC, USA
A*24:3021739-6232-5 ,KX622666,Yuxin Yin, Los Angeles, USA
B*07:02:03NT01424,KX583631,Carolyn K Hurley, Washington DC, USA
B*15:04:04261881,KX430223,Raquel Fabreti, Belo Horizonte, Brazil
B*27:13RDP-CH6-57,KX686520,Carolyn K Hurley, Washington DC, USA
B*40:3201473-7874-9 ,KX622685,Yuxin Yin, Los Angeles, USA
B*40:331221572,KX430224,Raquel Fabreti, Belo Horizonte, Brazil
B*40:335HG00012239,KX578053,Histogenetics, Ossining, USA
B*44:2211479-7990-0 ,KX622684,Yuxin Yin, Los Angeles, USA
B*52:39JRC-D79-074,LC171874,Masahiro Satake, Tokyo, Japan
C*04:238015876306,LT595714,Dominique Masson, La Tronche, France
C*06:02:07NT01413,KX583641,Carolyn K Hurley, Washington DC, USA
C*07:02:01:03JY,AJ293016,Ann-Margaret Little, Glasgow, United Kingdom
C*08:53NT01414,KX583640,Carolyn K Hurley, Washington DC, USA
C*12:03:06NT01415,KX583639,Carolyn K Hurley, Washington DC, USA
C*12:12NT01416,KX583638,Carolyn K Hurley, Washington DC, USA
C*12:188DKMS-LSL-C-680,LN912947,DKMS Life Sciences Lab, Dresden, Germany
DPA1*01:03:01:01TUBO,KU239959,Konstantinos Barsakis, Palo Alto, USA
DPA1*02:01:01:02DAN723,KU239963,Konstantinos Barsakis, Palo Alto, USA
DPA1*02:01:02KLO,KU239965,Konstantinos Barsakis, Palo Alto, USA
DPA1*02:02:02FH44,KU239967,Konstantinos Barsakis, Palo Alto, USA
DPA1*03:01SHJO,KU239968,Konstantinos Barsakis, Palo Alto, USA
DPA1*04:01CHI007,KU239969,Konstantinos Barsakis, Palo Alto, USA
DQB1*04:01:01FH44,KU319365,Konstantinos Barsakis, Palo Alto, USA
DQB1*05:971742-4047-3 ,KX622651,Yuxin Yin, Los Angeles, USA
DQB1*05:1031742-1894-1 ,KX622649,Yuxin Yin, Los Angeles, USA
DQB1*06:79:01BY01307,KX686513,Carolyn K Hurley, Washington DC, USA
DRB1*01:74241282,KX646400,Raquel Fabreti, Belo Horizonte, Brazil
DRB1*03:1241671-2966-7 ,KX622658,Yuxin Yin, Los Angeles, USA
DRB1*04:02:01FKD1040201,KX687269,Konstantinos Barsakis, Palo Alto, USA
DRB1*04:03:121720-9672-9 ,KX622654,Yuxin Yin, Los Angeles, USA
DRB1*04:04:01FKD1040401E2,KX687271,Konstantinos Barsakis, Palo Alto, USA
DRB1*04:05:01FH44,KX687272,Konstantinos Barsakis, Palo Alto, USA
DRB1*04:06:01ISH3,KX687243,Konstantinos Barsakis, Palo Alto, USA
DRB1*04:16BEL5GB,KX687245,Konstantinos Barsakis, Palo Alto, USA
DRB1*08:01:01BM9,KX687246,Konstantinos Barsakis, Palo Alto, USA
DRB1*08:04:01FK080401,KX687248,Konstantinos Barsakis, Palo Alto, USA
DRB1*08:14FH44,KX687249,Konstantinos Barsakis, Palo Alto, USA
DRB1*09:01:02280599,KX687274,Konstantinos Barsakis, Palo Alto, USA
DRB1*11:09BEL7MON,KX687251,Konstantinos Barsakis, Palo Alto, USA
DRB1*12:02:01FK120201,KX687254,Konstantinos Barsakis, Palo Alto, USA
DRB1*13:03:01FKD1130301E1,KX687275,Konstantinos Barsakis, Palo Alto, USA
DRB1*13:05:01FKD1130501E1,KX687276,Konstantinos Barsakis, Palo Alto, USA
DRB1*14:02:01DAN723,KX687277,Konstantinos Barsakis, Palo Alto, USA
DRB1*14:06:01FKD1140601E1,KX687278,Konstantinos Barsakis, Palo Alto, USA
DRB1*14:09BUR,E,KX687261,Konstantinos Barsakis, Palo Alto, USA
DRB1*14:14IHL, AD036,KX687262,Konstantinos Barsakis, Palo Alto, USA
DRB1*15:02:02THA,KX687263,Konstantinos Barsakis, Palo Alto, USA
DRB1*16:01:01CHA,AJ,KX687264,Konstantinos Barsakis, Palo Alto, USA
DRB1*16:04FORE,KX687283,Konstantinos Barsakis, Palo Alto, USA

Table 3: Recently Published Sequences


Sequence References
A*23:01:19(3)
A*24:325(4)
B*13:69(5)
B*38:15(6)
B*44:237N(7)
B*58:80(8)
DRB1*09:28(9)
DRB1*16:39(10)

References:

  1. Marsh SGE, Albert ED, Bodmer WF, et al. Nomenclature for factors of the HLA system, 2010. Tissue Antigens 2010 75: 291-455.
  2. Robinson J, Halliwell JA, Hayhurst JD, Flicek P, Parham P, Marsh SGE. The IPD and IMGT/HLA database: allele variant databases. Nucleic Acids Res 2015: 43: 423-31.
  3. Sidahmed AME, Anderson J, Stamm L, et al.HLA-A*23:01:19, a novel variant of HLA-A*23:01, discovered in a West African stem cell transplantation patient. HLA 2016 88: 120-121.
  4. Gao SQ, Zhong YP, Zhang H, et al.A novel HLA-A*24 allele, HLA-A*24:325, identified in a Chinese individual. HLA 2016 88: 121-122.
  5. Zheng JW, Xie JH, Sun Y, et al.A novel HLA-B allele, HLA-B*13:69. HLA 2016 88: 122-123.
  6. Yang KL, Hung JH, Lin PY.A novel HLA-B*38 variant, B*38:15, detected in a Taiwanese individual HLA 2016 88: 125-126.
  7. Qiang W, Zhu YY, Shen G, et al.Identification of a novel HLA-B*44 allele, HLA-B*44:237N, in a Chinese individual. HLA 2016 88: 126-127.
  8. Yang KL, HUng JH, Lin PY.Discovery of HLA-B*58:80, a novel HLA-B*58 variant, in a Taiwanese bone marrow donor. HLA 2016 88: 127-129.
  9. Yang KL, Hung JH, Lin PY.Discovery of a novel HLA-DRB1*09 variant, HLA-DRB1*09:28, in a Taiwanese individual. HLA 2016 88: 129-130.
  10. Yang KL, Hung JH, Lin PY.HLA-DRB1*16:39, a novel HLA-DRB1*16 variant, discovered in a Taiwanese bone marrow hematopoietic stem cell donor. HLA 2016 88: 124-125.