Nomenclature for Factors of the HLA System
July 2014
Compiled by Steven G. E. Marsh for the WHO Nomenclature Committee for Factors of the HLA System.
Published In:
- Tissue Antigens (2014) 84:598-603
- Human Immunology (2014) 75:1276-1279
- International Journal of Immunogenetics (2014) 41:530-534
The following sequences have been submitted to the Nomenclature Committee since the June 2014 nomenclature update and, following agreed policy, have been assigned official allele designations. Full details of all sequences will be published in a forthcoming report.
Below are listed the newly assigned sequences (Table 1) and confirmations of previously reported sequences (Table 2). The accession number of each sequence is given and these can be used to retrieve the sequence files from the EMBL, GenBank or DDBJ data libraries. Although accession numbers have been assigned by the data-libraries and most sequences are already available, there is still the possibility that an author may not yet have allowed the sequence to be released; in such a case you will have to contact the submitting author directly. Additional information pertaining to new sequences is often included in the publications describing these alleles; a listing of recent publications that describe new HLA sequences is given in Table 3.
All new and confirmatory sequences should now be submitted directly to the WHO Nomenclature Committee for Factors of the HLA System via the IMGT/HLA Database using the sequence submission tool provided (2). The IMGT/HLA Database may be accessed via the world wide web at : http://ww.ebi.ac.uk/imgt/hla/.
Table 1: New Sequences
| Sequence | Cell identification | Accession number | Submitting author |
|---|---|---|---|
| A*01:162N | HG00008695 | KJ819833 | Histogenetics, Ossining, USA |
| A*01:163 | HG00008699 | KJ906524 | Histogenetics, Ossining, USA |
| A*02:01:113 | HG00008702 | KJ923255 | Histogenetics, Ossining, USA |
| A*02:01:114 | HG00008703 | KJ956730 | Histogenetics, Ossining, USA |
| A*02:01:115 | HG00008707 | KJ923257 | Histogenetics, Ossining, USA |
| A*02:02:02 | HG00008708, HG00008709, HG00008710, HG00008711 | KJ923261, KJ956738, KJ956739, KJ956740 | Histogenetics, Ossining, USA |
| A*02:289:02 | HG00008706 | KJ956733 | Histogenetics, Ossining, USA |
| A*02:539 | CBRA59136263827 | LK391505 | Dominique Masson, La Tronche, France |
| A*02:540N | HG00008705 | KM012012 | Histogenetics, Ossining, USA |
| A*03:201 | HG00008694 | KJ956736 | Histogenetics, Ossining, USA |
| A*03:202 | HG00008697 | KJ906544 | Histogenetics, Ossining, USA |
| A*03:203 | HG00008698 | KM012013 | Histogenetics, Ossining, USA |
| A*11:207 | HG00008690 | KJ956734 | Histogenetics, Ossining, USA |
| A*23:67 | HG00008693 | KM012015 | Histogenetics, Ossining, USA |
| A*29:72 | DKMS-LSL-A-127 | LK021944 | DKMS Life Sciences Lab, Dresden, Germany |
| A*29:73 | DKMS-LSL-A-172 | LK021991 | DKMS Life Sciences Lab, Dresden, Germany |
| A*29:74 | HG00008688 | KJ956731 | Histogenetics, Ossining, USA |
| A*30:91 | HG00008692 | KJ923258 | Histogenetics, Ossining, USA |
| A*31:01:23 | DKMS-LSL-A-179 | LK021998 | DKMS Life Sciences Lab, Dresden, Germany |
| A*32:01:18 | HG00008696 | KJ923262 | Histogenetics, Ossining, USA |
| A*32:01:19 | HG00008700 | KJ906525 | Histogenetics, Ossining, USA |
| A*32:01:20 | HG00008701 | KJ956729 | Histogenetics, Ossining, USA |
| A*33:94 | HG00008687 | KJ923254 | Histogenetics, Ossining, USA |
| A*68:01:28 | HG00008691 | KJ956737 | Histogenetics, Ossining, USA |
| A*68:02:08 | HG00008689 | KJ923259 | Histogenetics, Ossining, USA |
| B*07:02:44 | HG00008762 | KJ819871 | Histogenetics, Ossining, USA |
| B*07:05:08 | HG00008763 | KJ819861 | Histogenetics, Ossining, USA |
| B*07:228 | HG00008760 | KJ923274 | Histogenetics, Ossining, USA |
| B*07:229 | HG00008761 | KJ906527 | Histogenetics, Ossining, USA |
| B*07:230 | HG00008764 | KJ819859 | Histogenetics, Ossining, USA |
| B*07:231N | HG00008765 | KM012016 | Histogenetics, Ossining, USA |
| B*07:232 | HG00008766 | KJ819857 | Histogenetics, Ossining, USA |
| B*08:135 | HG00008720 | KJ906545 | Histogenetics, Ossining, USA |
| B*08:136 | HG00008721 | KJ749714 | Histogenetics, Ossining, USA |
| B*13:01:09 | HG00008723 | KJ749725 | Histogenetics, Ossining, USA |
| B*14:01:06 | HG00008768 | KJ819860 | Histogenetics, Ossining, USA |
| B*14:41N | 0188-3540-5_NMDP | KM009989 | Carmen Alaez, Mexico City, Mexico |
| B*14:42 | HG00008758 | KJ923276 | Histogenetics, Ossining, USA |
| B*15:10:04 | HG00008751 | KJ749720 | Histogenetics, Ossining, USA |
| B*15:330 | HG00008738 | KJ923272 | Histogenetics, Ossining, USA |
| B*15:331 | HG00008742 | KJ906531 | Histogenetics, Ossining, USA |
| B*15:332 | HG00008743 | KJ819854 | Histogenetics, Ossining, USA |
| B*15:333 | HG00008744 | KJ906546 | Histogenetics, Ossining, USA |
| B*15:334 | HG00008745 | KJ819858 | Histogenetics, Ossining, USA |
| B*15:335 | HG00008752 | KJ956743 | Histogenetics, Ossining, USA |
| B*27:05:29 | HG00008717 | KJ749775 | Histogenetics, Ossining, USA |
| B*27:05:30 | HG00008718 | KJ956745 | Histogenetics, Ossining, USA |
| B*35:01:43 | HG00008733 | KJ956742 | Histogenetics, Ossining, USA |
| B*35:02:07 | HG00008732 | KJ956741 | Histogenetics, Ossining, USA |
| B*35:03:18 | HG00008729 | KJ749717 | Histogenetics, Ossining, USA |
| B*35:15:02 | DKMS-LSL-B-186 | LK022005 | DKMS Life Sciences Lab, Dresden, Germany |
| B*35:261 | BY01228 | KJ912624 | Carolyn K Hurley, Washington DC, USA |
| B*35:262 | HG00008731 | KJ923271 | Histogenetics, Ossining, USA |
| B*35:263 | HG00008769 | KJ923265 | Histogenetics, Ossining, USA |
| B*37:53 | HG00008719 | KJ749718 | Histogenetics, Ossining, USA |
| B*39:01:22 | HG00008759 | KJ906532 | Histogenetics, Ossining, USA |
| B*39:98 | HG00008767 | KJ819863 | Histogenetics, Ossining, USA |
| B*40:114:02 | BY01227 | KJ912626 | Carolyn K Hurley, Washington DC, USA |
| B*40:296 | HG00008716 | KJ906549 | Histogenetics, Ossining, USA |
| B*44:205 | HG00008722 | KJ749719 | Histogenetics, Ossining, USA |
| B*44:206 | HG00008724 | KM012017 | Histogenetics, Ossining, USA |
| B*44:207 | HG00008725 | KJ906528 | Histogenetics, Ossining, USA |
| B*44:208 | HG00008726 | KJ906547 | Histogenetics, Ossining, USA |
| B*44:209 | HG00008727 | KJ923263 | Histogenetics, Ossining, USA |
| B*44:210 | HG00008728 | KJ923270 | Histogenetics, Ossining, USA |
| B*46:61 | HG00008739 | KJ749721 | Histogenetics, Ossining, USA |
| B*46:62 | HG00008740 | KJ956744 | Histogenetics, Ossining, USA |
| B*46:63 | HG00008741 | KJ923273 | Histogenetics, Ossining, USA |
| B*49:01:07 | HG00008714 | KJ906530 | Histogenetics, Ossining, USA |
| B*49:32 | DKMS-LSL-B-203 | LK022022 | DKMS Life Sciences Lab, Dresden, Germany |
| B*49:33 | HG00008713 | KJ749776 | Histogenetics, Ossining, USA |
| B*51:92:02 | HG00008756 | KJ749715 | Histogenetics, Ossining, USA |
| B*51:180 | HG00008730 | KJ749716 | Histogenetics, Ossining, USA |
| B*51:181 | HG00008754 | KJ923266 | Histogenetics, Ossining, USA |
| B*51:182 | HG00008755 | KJ749713 | Histogenetics, Ossining, USA |
| B*51:183 | HG00008757 | KJ819865 | Histogenetics, Ossining, USA |
| B*52:01:21 | HG00008735 | KJ906548 | Histogenetics, Ossining, USA |
| B*53:38 | HG00008734 | KJ923264 | Histogenetics, Ossining, USA |
| B*55:01:13 | HG00008748 | KJ749724 | Histogenetics, Ossining, USA |
| B*55:69 | HG00008750 | KJ923275 | Histogenetics, Ossining, USA |
| B*56:01:08 | HG00008749 | KJ749722 | Histogenetics, Ossining, USA |
| B*57:14:02 | HG00008746 | KJ819862 | Histogenetics, Ossining, USA |
| B*57:73 | HG00008715 | KJ923269 | Histogenetics, Ossining, USA |
| B*57:74 | HG00008747 | KJ906529 | Histogenetics, Ossining, USA |
| B*58:62 | HG00008736 | KJ819853 | Histogenetics, Ossining, USA |
| B*78:08 | HG00008753 | KJ749723 | Histogenetics, Ossining, USA |
| C*02:02:28 | HG00008646 | KJ819888 | Histogenetics, Ossining, USA |
| C*02:02:29 | HG00008795 | KM012019 | Histogenetics, Ossining, USA |
| C*02:87 | DKMS-LSL-C-144 | LK021962 | DKMS Life Sciences Lab, Dresden, Germany |
| C*02:88 | HG00008645 | KJ819879 | Histogenetics, Ossining, USA |
| C*03:263 | HG00008652 | KJ749733 | Histogenetics, Ossining, USA |
| C*03:264 | HG00008786 | KJ956754 | Histogenetics, Ossining, USA |
| C*03:265N | HG00008799 | KJ906533 | Histogenetics, Ossining, USA |
| C*03:266 | HG00008800 | KJ819876 | Histogenetics, Ossining, USA |
| C*04:01:63 | HG00008654 | KJ819892 | Histogenetics, Ossining, USA |
| C*04:01:64 | HG00008805 | KJ956750 | Histogenetics, Ossining, USA |
| C*04:178 | HG00008656 | KJ819891 | Histogenetics, Ossining, USA |
| C*04:179 | HG00008802 | KJ923279 | Histogenetics, Ossining, USA |
| C*04:180 | HG00008803 | KJ956746 | Histogenetics, Ossining, USA |
| C*04:181 | HG00008806 | KM012023 | Histogenetics, Ossining, USA |
| C*05:01:28 | HG00008638 | KJ819890 | Histogenetics, Ossining, USA |
| C*05:01:29 | HG00008653 | KJ819883 | Histogenetics, Ossining, USA |
| C*05:01:30 | HG00008787 | KJ749783 | Histogenetics, Ossining, USA |
| C*05:104 | HG00008639 | KJ749730 | Histogenetics, Ossining, USA |
| C*06:127:02 | HG00008772 | KM012020 | Histogenetics, Ossining, USA |
| C*06:138 | HG00008771 | KJ956748 | Histogenetics, Ossining, USA |
| C*06:139 | HG00008774 | KM012018 | Histogenetics, Ossining, USA |
| C*06:140 | HG00008775 | KJ819878 | Histogenetics, Ossining, USA |
| C*06:141 | HG00008776 | KJ906551 | Histogenetics, Ossining, USA |
| C*07:131:02 | HG00008807 | KJ749784 | Histogenetics, Ossining, USA |
| C*07:393N | D227885 | LK021326 | Edward KL Yang, Hualien, Taiwan |
| C*07:394 | HG00008777 | KJ956751 | Histogenetics, Ossining, USA |
| C*07:395 | HG00008778 | KJ923280 | Histogenetics, Ossining, USA |
| C*07:396 | HG00008780 | KM012024 | Histogenetics, Ossining, USA |
| C*07:397 | HG00008781 | KJ923285 | Histogenetics, Ossining, USA |
| C*07:398 | HG00008782 | KJ906552 | Histogenetics, Ossining, USA |
| C*07:399 | HG00008783 | KJ956747 | Histogenetics, Ossining, USA |
| C*07:400 | HG00008784 | KJ923282 | Histogenetics, Ossining, USA |
| C*07:401 | HG00008785 | KJ923283 | Histogenetics, Ossining, USA |
| C*08:01:18 | HG00008788 | KJ956749 | Histogenetics, Ossining, USA |
| C*12:02:13 | HG00008791 | KM012021 | Histogenetics, Ossining, USA |
| C*12:130 | HG00008640 | KJ749736 | Histogenetics, Ossining, USA |
| C*12:131 | HG00008641 | KJ749785 | Histogenetics, Ossining, USA |
| C*12:132 | HG00008643 | KJ819884 | Histogenetics, Ossining, USA |
| C*12:133 | HG00008789 | KJ906553 | Histogenetics, Ossining, USA |
| C*12:134 | HG00008790 | KJ819874 | Histogenetics, Ossining, USA |
| C*14:02:18 | HG00008655 | KJ749786 | Histogenetics, Ossining, USA |
| C*14:68 | HG00008801 | KJ923278 | Histogenetics, Ossining, USA |
| C*15:02:20 | HG00008648, HG00008649, HG00008650 | KJ819881, KJ749731, KJ749729 | Histogenetics, Ossining, USA |
| C*15:97 | D10056013 | KF175506 | Jinghui Qu, Shenzhen, China |
| C*15:98 | HG00008647 | KJ749737 | Histogenetics, Ossining, USA |
| C*15:99 | HG00008651 | KJ749727 | Histogenetics, Ossining, USA |
| C*15:100 | HG00008796 | KJ923277 | Histogenetics, Ossining, USA |
| C*15:101 | HG00008797 | KJ923281 | Histogenetics, Ossining, USA |
| C*16:01:19 | HG00008644 | KJ819885 | Histogenetics, Ossining, USA |
| C*16:76 | HG00008642 | KJ819894 | Histogenetics, Ossining, USA |
| C*17:26 | HG00008792 | KJ923284 | Histogenetics, Ossining, USA |
| C*17:27N | HG00008793 | KJ906550 | Histogenetics, Ossining, USA |
| C*18:08 | HG00008773 | KJ906554 | Histogenetics, Ossining, USA |
| C*18:09 | HG00008794 | KJ906534 | Histogenetics, Ossining, USA |
| DRB1*01:64 | HG00008675 | KJ819905 | Histogenetics, Ossining, USA |
| DRB1*03:113 | HG00008670 | KJ819906 | Histogenetics, Ossining, USA |
| DRB1*03:114 | HG00008827 | KJ906537 | Histogenetics, Ossining, USA |
| DRB1*04:182 | HG00008668 | KJ749792 | Histogenetics, Ossining, USA |
| DRB1*04:183 | HG00008669 | KJ749750 | Histogenetics, Ossining, USA |
| DRB1*04:184 | HG00008826 | KJ923299 | Histogenetics, Ossining, USA |
| DRB1*11:168 | HG00008673 | KJ819907 | Histogenetics, Ossining, USA |
| DRB1*13:188 | HG00008828 | KJ819910 | Histogenetics, Ossining, USA |
| DRB1*13:189 | HG00008829 | KM012026 | Histogenetics, Ossining, USA |
| DRB1*14:158 | HG00008671, HG00008672 | KJ819909, KJ819908 | Histogenetics, Ossining, USA |
| DRB1*14:159 | HG00008674 | KJ819912 | Histogenetics, Ossining, USA |
| DRB1*15:01:25 | HG00008822 | KJ956762 | Histogenetics, Ossining, USA |
| DRB1*15:01:26 | HG00008823 | KJ923297 | Histogenetics, Ossining, USA |
| DRB1*15:06:03 | HG00008821 | KJ923298 | Histogenetics, Ossining, USA |
| DRB1*15:113N | HG00008824, HG00008825 | KJ906556, KJ906557 | Histogenetics, Ossining, USA |
| DQB1*02:01:18 | HG00008662 | KJ749748 | Histogenetics, Ossining, USA |
| DQB1*02:01:19 | HG00008811 | KJ923287 | Histogenetics, Ossining, USA |
| DQB1*02:14:02 | HG00008810 | KJ956755 | Histogenetics, Ossining, USA |
| DQB1*02:42 | HG00008661 | KJ819895 | Histogenetics, Ossining, USA |
| DQB1*03:01:25 | HG00008808 | KJ906535 | Histogenetics, Ossining, USA |
| DQB1*03:151 | HG00008659 | KJ819899 | Histogenetics, Ossining, USA |
| DQB1*03:152 | HG00008660 | KJ749790 | Histogenetics, Ossining, USA |
| DQB1*03:153 | HG00008664 | KJ749744 | Histogenetics, Ossining, USA |
| DQB1*03:154 | HG00008809 | KM012025 | Histogenetics, Ossining, USA |
| DQB1*03:155 | HG00008812 | KJ923295 | Histogenetics, Ossining, USA |
| DQB1*03:156 | HG00008813 | KJ923288 | Histogenetics, Ossining, USA |
| DQB1*03:157 | HG00008816 | KJ956761 | Histogenetics, Ossining, USA |
| DQB1*05:01:18 | HG00008658 | KJ819896 | Histogenetics, Ossining, USA |
| DQB1*05:79 | HG00008657 | KJ819902 | Histogenetics, Ossining, USA |
| DQB1*05:80 | HG00008665 | KJ749743 | Histogenetics, Ossining, USA |
| DQB1*05:81 | HG00008666 | KJ819900 | Histogenetics, Ossining, USA |
| DQB1*05:82 | HG00008814 | KJ956758 | Histogenetics, Ossining, USA |
| DQB1*05:83 | HG00008815 | KJ906536 | Histogenetics, Ossining, USA |
| DQB1*06:02:21 | HG00008667 | KJ819898 | Histogenetics, Ossining, USA |
| DQB1*06:160 | HG00008663 | KJ749742 | Histogenetics, Ossining, USA |
| DQB1*06:161 | HG00008817 | KJ819904 | Histogenetics, Ossining, USA |
| DQB1*06:162 | HG00008818 | KJ923289 | Histogenetics, Ossining, USA |
| DQB1*06:163 | HG00008819, HG00008820 |
KJ956757, KJ923296 | Histogenetics, Ossining, USA |
| DPB1*02:01:14 | HG00008677 | KJ819922 | Histogenetics, Ossining, USA |
| DPB1*02:01:15 | HG00008842, HG00008843 |
KJ923305, KJ819918 | Histogenetics, Ossining, USA |
| DPB1*03:01:04 | HG00008856 | KJ923311 | Histogenetics, Ossining, USA |
| DPB1*04:01:24 | HG00008846, HG00008847 |
KJ923302, KJ923303 | Histogenetics, Ossining, USA |
| DPB1*04:02:07 | HG00008678 | KJ749763 | Histogenetics, Ossining, USA |
| DPB1*05:01:03 | HG00008835 | KJ819920 | Histogenetics, Ossining, USA |
| DPB1*05:01:04 | HG00008836 | KM012036 | Histogenetics, Ossining, USA |
| DPB1*354:01:02 | HG00008833 | KJ749756 | Histogenetics, Ossining, USA |
| DPB1*364:01 | HG00008676 | KJ819928 | Histogenetics, Ossining, USA |
| DPB1*365:01 | HG00008679 | KJ585785 | Histogenetics, Ossining, USA |
| DPB1*366:01 | HG00008680 | KJ819931 | Histogenetics, Ossining, USA |
| DPB1*367:01 | HG00008681 | KJ819923 | Histogenetics, Ossining, USA |
| DPB1*368:01 | HG00008682 | KJ819924 | Histogenetics, Ossining, USA |
| DPB1*369:01 | HG00008683 | KJ819921 | Histogenetics, Ossining, USA |
| DPB1*370:01 | HG00008684 | KJ749795 | Histogenetics, Ossining, USA |
| DPB1*371:01 | HG00008685 | KJ749759 | Histogenetics, Ossining, USA |
| DPB1*372:01 | HG00008830 | KJ906539 | Histogenetics, Ossining, USA |
| DPB1*373:01 | HG00008831 | KJ956778 | Histogenetics, Ossining, USA |
| DPB1*374:01 | HG00008832 | KJ956768 | Histogenetics, Ossining, USA |
| DPB1*375:01 | HG00008834 | KJ956771 | Histogenetics, Ossining, USA |
| DPB1*376:01 | HG00008837 | KJ749758 | Histogenetics, Ossining, USA |
| DPB1*377:01 | HG00008838 | KJ956767 | Histogenetics, Ossining, USA |
| DPB1*378:01 | HG00008839 | KM012039 | Histogenetics, Ossining, USA |
| DPB1*379:01 | HG00008840 | KJ906538 | Histogenetics, Ossining, USA |
| DPB1*380:01 | HG00008841 | KJ923312 | Histogenetics, Ossining, USA |
| DPB1*381:01 | HG00008844 | KJ956775 | Histogenetics, Ossining, USA |
| DPB1*382:01N | HG00008845 | KM012027 | Histogenetics, Ossining, USA |
| DPB1*383:01 | HG00008848, HG00008849 |
KJ819917, KJ819916 | Histogenetics, Ossining, USA |
| DPB1*384:01 | HG00008850 | KJ819915 | Histogenetics, Ossining, USA |
| DPB1*385:01 | HG00008851 | KJ923316 | Histogenetics, Ossining, USA |
| DPB1*386:01 | HG00008852 | KJ923320 | Histogenetics, Ossining, USA |
| DPB1*387:01 | HG00008853 | KJ923306 | Histogenetics, Ossining, USA |
| DPB1*388:01 | HG00008854 | KJ923315 | Histogenetics, Ossining, USA |
| DPB1*389:01 | HG00008855 | KJ956769 | Histogenetics, Ossining, USA |
| DPB1*390:01 | HG00008857 | KJ819914 | Histogenetics, Ossining, USA |
| DPB1*391:01 | HG00008858 | KJ923318 | Histogenetics, Ossining, USA |
| DPB1*392:01 | HG00008859 | KJ906558 | Histogenetics, Ossining, USA |
Table 2: Confirmatory Sequences
| Sequence | Cell identification | Accession number | Submitting author |
|---|---|---|---|
| A*02:01:103 | HG00008712 | KM012014 | Histogenetics, Ossining, USA |
| A*11:196 | 368481 | LK021327 | Edward KL Yang, Hualien, Taiwan |
| A*11:199 | DKMS-LSL-A-129 | LK021946 | DKMS Life Sciences Lab, Dresden, Germany |
| B*51:174 | 62135758884 | LK391506 | Dominique Masson, La Tronche, France |
| C*04:01:62 | HG00008804 | KJ749781 | Histogenetics, Ossining, USA |
| C*07:386 | HG00008779 | KJ819877 | Histogenetics, Ossining, USA |
| C*15:85 | HG00008798 | KJ819875 | Histogenetics, Ossining, USA |
| DRB1*03:15 | 31571 | LK391773 | Marcel GJ Tilanus, Maastricht, The Netherlands |
| DRB1*08:16 | 30063 | LK391772 | Marcel GJ Tilanus, Maastricht, The Netherlands |
| DRB1*08:35 | 39200 | LK391783 | Marcel GJ Tilanus, Maastricht, The Netherlands |
| DRB1*13:06 | 32795 | LK391775 | Marcel GJ Tilanus, Maastricht, The Netherlands |
| DRB1*13:11:01 | 31704 | LK391774 | Marcel GJ Tilanus, Maastricht, The Netherlands |
| DRB1*14:06:01 | 33401 | LK391776 | Marcel GJ Tilanus, Maastricht, The Netherlands |
| DRB1*14:21 | 23482 | LK391771 | Marcel GJ Tilanus, Maastricht, The Netherlands |
| DQB1*02:14:01 | HIP04607 | KM017969 | Anajane Smith, Seattle, USA |
Table 3: Recently Published Sequences
| Sequence | References |
|---|---|
| A*02:433 | (3) |
| A*02:434 | (3) |
| DRB1*11:150 | (4) |
| DRB1*14:145 | (4) |
References:
- Marsh SGE, Albert ED, Bodmer WF, et al. Nomenclature for Factors of the HLA system, 2010. Tissue Antigens. 2010; 75:291-455.
- Robinson J, Halliwell JA, McWilliam H, Lopez R, Parham P, Marsh SGE. The IMGT/HLA database. Nucleic Acids Research. 2013; 41: D1222-7.
- Fakhoury HA, Jawdat D, Alaskar AS, Al Jumah M, Cereb N, Hajeer AH. Two novel alleles HLA-A*02:433 and HLA-A*02:434 identified in Saudi bone marrow donors using sequence-based typing. International Journal of Immunogenetics. 2014; 41: 338-9.
- Fakhoury HA, Cereb N, Jawdat D, Al Jumah M, Alaskar AS, Hajeer AH. Two novel alleles HLA-DRB1*11:150 and HLA-DRB1*14:145 identified in Saudi individuals. International Journal of Immunogenetics. 2014; 41: 340-1.
