Nomenclature for Factors of the HLA System
February 2012
Compiled by Steven G. E. Marsh for the WHO Nomenclature Committee for Factors of the HLA System.
Published In:
- Tissue Antigens. 2012 80:72-7
- Human Immunology. 2012 73:767-71
- International Journal of Immunogenetics. 2012 39:365-9
The following sequences have been submitted to the Nomenclature Committee since the January 2012 nomenclature update and, following agreed policy, have been assigned official allele designations. Full details of all sequences will be published in a forthcoming report.
Below are listed the newly assigned sequences (Table 1) and confirmations of previously reported sequences (Table 2). The accession number of each sequence is given and these can be used to retrieve the sequence files from the EMBL, GenBank or DDBJ data libraries. Although accession numbers have been assigned by the data-libraries and most sequences are already available, there is still the possibility that an author may not yet have allowed the sequence to be released; in such a case you will have to contact the submitting author directly. Additional information pertaining to new sequences is often included in the publications describing these alleles; a listing of recent publications that describe new HLA sequences is given in Table 3.
All new and confirmatory sequences should now be submitted directly to the WHO Nomenclature Committee for Factors of the HLA System via the IMGT/HLA Database using the sequence submission tool provided (2). The IMGT/HLA Database may be accessed via the world wide web at : http://ww.ebi.ac.uk/imgt/hla/.
Table 1: New Sequences
Sequence | Cell identification | Accession number | Submitting author |
---|---|---|---|
A*01:01:44 | HG00002793, HG00002794 | JN604843, JN604844 | Histogenetics, Ossining, USA |
A*01:01:45 | HG00002963 | JN561712 | Histogenetics, Ossining, USA |
A*01:105 | HG00002760 | JN124029 | Histogenetics, Ossining, USA |
A*01:106 | HG00002770 | JN604842 | Histogenetics, Ossining, USA |
A*01:107 | WEIMano | HE660039 | Sylvie Tourne, Strasbourg, France |
A*02:01:67 | HG00002804 | JN124079 | Histogenetics, Ossining, USA |
A*02:01:68 | HG00002817 | JN124031 | Histogenetics, Ossining, USA |
A*02:01:69 | HG00002824 | JN123973 | Histogenetics, Ossining, USA |
A*02:01:70 | HG00002827, HG00002828 | JN124101, JN561711 | Histogenetics, Ossining, USA |
A*02:01:71 | HG00002832 | JN123752 | Histogenetics, Ossining, USA |
A*02:01:72 | 55486 | JQ429790 | Jianping Li, Shenyang, Liaoning, China |
A*02:01:73 | 93081372 | JQ645967 | Daniel Fuerst, Ulm, Germany |
A*02:06:11 | HG00002798, HG00002799 | JN604840, JN123996 | Histogenetics, Ossining, USA |
A*02:344 | HG00002797 | JN406393 | Histogenetics, Ossining, USA |
A*02:345 | HG00002805 | JN584490 | Histogenetics, Ossining, USA |
A*02:346 | HG00002806 | JN406433 | Histogenetics, Ossining, USA |
A*02:347 | HG00002807 | JN124050 | Histogenetics, Ossining, USA |
A*02:348 | HG00002812 | JN124083 | Histogenetics, Ossining, USA |
A*02:349 | HG00002830 | JN124037 | Histogenetics, Ossining, USA |
A*02:350N | HG00002831 | JN119365 | Histogenetics, Ossining, USA |
A*02:351 | HG00002833, HG00002834 | JN124094, JN124076 | Histogenetics, Ossining, USA |
A*02:352 | HG00002840 | JN182958 | Histogenetics, Ossining, USA |
A*02:353 | HG00002969 | JN182964 | Histogenetics, Ossining, USA |
A*02:354 | HG00002973 | JN124127 | Histogenetics, Ossining, USA |
A*02:355 | 5381000333 | JN209961 | Chuanfu Zhu, Jinan, China |
A*02:356N | JRC-D01-092 | AB696719 | Kenji Tadokoro, Tokyo, Japan |
A*03:01:32 | HG00002778 | JN123998 | Histogenetics, Ossining, USA |
A*03:01:33 | HG00002786, HG00002787 | JN124088, JN124089 | Histogenetics, Ossining, USA |
A*03:137 | HG00002759 | JN124002 | Histogenetics, Ossining, USA |
A*03:138 | HG00002779 | JN124086 | Histogenetics, Ossining, USA |
A*03:139 | HG00002781 | JN182956 | Histogenetics, Ossining, USA |
A*03:140 | HG00002782 | JN124015 | Histogenetics, Ossining, USA |
A*03:141 | HG00002966 | JN406434 | Histogenetics, Ossining, USA |
A*03:142 | HG00002967 | JN584497 | Histogenetics, Ossining, USA |
A*11:01:38 | HG00002700 | JN124082 | Histogenetics, Ossining, USA |
A*11:01:39 | HG00002705, HG00002706 | JN418812, JN629077 | Histogenetics, Ossining, USA |
A*11:01:40 | HG00002957 | JN124113 | Histogenetics, Ossining, USA |
A*11:114 | HG00002698 | JN123992 | Histogenetics, Ossining, USA |
A*11:115N | HG00002699 | JN406502 | Histogenetics, Ossining, USA |
A*11:116 | HG00002704 | JN124093 | Histogenetics, Ossining, USA |
A*11:117 | HG00002709 | JN124100 | Histogenetics, Ossining, USA |
A*11:118 | HG00002710 | JN124005 | Histogenetics, Ossining, USA |
A*11:119 | HG00002953 | JN124109 | Histogenetics, Ossining, USA |
A*11:120 | 44461 | JQ365626 | Jianping Li, Shenyang, Liaoning, China |
A*23:01:09 | HG00002749 | JN644950 | Histogenetics, Ossining, USA |
A*23:51 | 87738 | JQ661498 | Maria Elisa Moraes, Rio de Janeiro, Brazil |
A*24:02:55 | GDY-1219 | JN590043 | Guolong Yu, Guanghzhou City, China |
A*24:02:56 | 55309 | JQ410444 | Jianping Li, Shenyang, Liaoning, China |
A*24:192 | HG00002733 | JN406388 | Histogenetics, Ossining, USA |
A*24:193 | HG00002735 | JN644953 | Histogenetics, Ossining, USA |
A*24:194 | HG00002740 | JN629076 | Histogenetics, Ossining, USA |
A*24:195 | HG00002741 | JN242427 | Histogenetics, Ossining, USA |
A*24:196 | HG00002742 | JN123987 | Histogenetics, Ossining, USA |
A*24:197 | HG00002744 | JN124052 | Histogenetics, Ossining, USA |
A*24:198 | 55031 | JQ392563 | Jianping Li, Shenyang, Liaoning, China |
A*24:199 | An 209308 | HE614129 | Francesca Poli, Milano, Italy |
A*25:01:06 | HG00002954 | JN584502 | Histogenetics, Ossining, USA |
A*25:01:07 | HG00002801 | JN124096 | Histogenetics, Ossining, USA |
A*26:01:23 | HG00002719 | JN418807 | Histogenetics, Ossining, USA |
A*26:01:24 | HG00002721 | JN182952 | Histogenetics, Ossining, USA |
A*26:73 | HG00002955 | JN124115 | Histogenetics, Ossining, USA |
A*26:74 | 5371000179 | JN209960 | Chuanfu Zhu, Jinan, China |
A*29:02:11 | HG00002677 | JN205179 | Histogenetics, Ossining, USA |
A*29:33 | HG00002676 | JN205169 | Histogenetics, Ossining, USA |
A*29:34 | HG00002945 | JN561715 | Histogenetics, Ossining, USA |
A*29:35 | HG00002946, HG00002947 | JN124155, JN584499 | Histogenetics, Ossining, USA |
A*30:60 | HG00002723 | JN123991 | Histogenetics, Ossining, USA |
A*30:61 | HG00002724, HG00002725 | JN124039, JN205178 | Histogenetics, Ossining, USA |
A*30:62 | 55160 | JQ392564 | Jianping Li, Shenyang, Liaoning, China |
A*31:01:12 | HG00002675 | JN124013 | Histogenetics, Ossining, USA |
A*31:62 | VAUGuil | HE660040 | Sylvie Tourne, Strasbourg, France |
A*32:38 | HG00002789 | JN124022 | Histogenetics, Ossining, USA |
A*32:39 | HG00002791 | JN418804 | Histogenetics, Ossining, USA |
A*32:40 | HG00002796 | JN561708 | Histogenetics, Ossining, USA |
A*33:32:02 | HG00002669 | JN124067 | Histogenetics, Ossining, USA |
A*33:56 | HG00002667 | JN123994 | Histogenetics, Ossining, USA |
A*33:57 | HG00002670 | JN182957 | Histogenetics, Ossining, USA |
A*33:58 | HG00002673, HG00002674 | JN123974, JN123993 | Histogenetics, Ossining, USA |
A*34:02:03 | HG00002716 | JN205172 | Histogenetics, Ossining, USA |
A*68:01:12 | HG00002718 | JN561705 | Histogenetics, Ossining, USA |
A*68:01:13 | HG00002722 | JN124040 | Histogenetics, Ossining, USA |
A*68:01:14 | HG00002956 | JN119373 | Histogenetics, Ossining, USA |
A*68:01:15 | HG00002958 | JN561714 | Histogenetics, Ossining, USA |
A*68:02:05 | HG00002697 | JN418806 | Histogenetics, Ossining, USA |
A*68:86 | HG00002696 | JN182949 | Histogenetics, Ossining, USA |
A*68:87 | HG00002717 | JN629080 | Histogenetics, Ossining, USA |
A*68:88 | 96009895 | JQ645966 | Daniel Fuerst, Ulm, Germany |
B*07:02:31 | HG00002928 | JN124184 | Histogenetics, Ossining, USA |
B*07:02:32 | HG00003088 | JN604856 | Histogenetics, Ossining, USA |
B*07:149 | HG00002924, HG00002925, HG00002926 | JN124242, JN124243, JN124294 | Histogenetics, Ossining, USA |
B*07:150 | HG00002927 | JN124174 | Histogenetics, Ossining, USA |
B*07:151 | HG00002932 | JN124191 | Histogenetics, Ossining, USA |
B*07:152 | HG00003087 | JN124258 | Histogenetics, Ossining, USA |
B*07:153 | HG00003093 | JN182976 | Histogenetics, Ossining, USA |
B*07:154 | HG00003098 | JN406455 | Histogenetics, Ossining, USA |
B*08:01:17 | HG00002873 | JN406450 | Histogenetics, Ossining, USA |
B*08:86N | HG00002871 | JN406503 | Histogenetics, Ossining, USA |
B*08:87 | HG00002872 | JN124202 | Histogenetics, Ossining, USA |
B*08:88 | HG00002878 | JN124218 | Histogenetics, Ossining, USA |
B*13:53 | HG00003016 | JN205203 | Histogenetics, Ossining, USA |
B*13:54 | HG00003017 | JN205210 | Histogenetics, Ossining, USA |
B*14:30 | HG00002923 | JN561720 | Histogenetics, Ossining, USA |
B*14:31 | HG00003083 | JN124289 | Histogenetics, Ossining, USA |
B*14:32 | HG00003106 | JN124299 | Histogenetics, Ossining, USA |
B*15:01:28 | HG00003053 | JN406412 | Histogenetics, Ossining, USA |
B*15:47:02 | HG00002913 | JN124213 | Histogenetics, Ossining, USA |
B*15:240 | HG00002904 | JN124196 | Histogenetics, Ossining, USA |
B*15:241 | HG00002905 | JN124179 | Histogenetics, Ossining, USA |
B*15:242 | HG00002915 | JN124211 | Histogenetics, Ossining, USA |
B*15:243 | HG00003065 | JN604858 | Histogenetics, Ossining, USA |
B*15:244 | HG00003066 | JN205194 | Histogenetics, Ossining, USA |
B*18:01:01:02 | 18.1 | HM161847 | David Sayer, Perth, Australia |
B*18:01:15 | HG00002863 | JN182974 | Histogenetics, Ossining, USA |
B*18:01:16 | HG00003007 | JN182978 | Histogenetics, Ossining, USA |
B*18:69 | HG00002861 | JN561723 | Histogenetics, Ossining, USA |
B*18:70 | HG00002979 | JN119421 | Histogenetics, Ossining, USA |
B*27:05:18 | HG00002865, HG00002866 | JN124183, JN124244 | Histogenetics, Ossining, USA |
B*27:88 | HG00002857 | JN123810 | Histogenetics, Ossining, USA |
B*27:89 | HG00002995 | JN584513 | Histogenetics, Ossining, USA |
B*35:01:30 | HG00002893 | JN644964 | Histogenetics, Ossining, USA |
B*35:01:31 | HG00003043 | JN418824 | Histogenetics, Ossining, USA |
B*35:03:11 | HG00003045 | JN406457 | Histogenetics, Ossining, USA |
B*35:190 | HG00002894 | JN644966 | Histogenetics, Ossining, USA |
B*35:191 | HG00002897, HG00002898 | JN124192, JN406449 | Histogenetics, Ossining, USA |
B*35:192 | HG00002911 | JN205187 | Histogenetics, Ossining, USA |
B*35:193 | HG00003014 | JN406451 | Histogenetics, Ossining, USA |
B*35:194 | HG00003033 | JN205195 | Histogenetics, Ossining, USA |
B*35:195 | HG00003036 | JN182979 | Histogenetics, Ossining, USA |
B*35:196 | HG00003046 | JN124271 | Histogenetics, Ossining, USA |
B*35:197 | HG00003085 | JN124291 | Histogenetics, Ossining, USA |
B*35:198 | 29524 | JQ627828 | Maria Elisa Moraes, Rio de Janeiro, Brazil |
B*37:33N | HG00003001 | JN123788 | Histogenetics, Ossining, USA |
B*38:01:07 | HG00003115 | JN406408 | Histogenetics, Ossining, USA |
B*38:39 | HG00003084 | JN124283 | Histogenetics, Ossining, USA |
B*38:40 | HG00003086 | JN584506 | Histogenetics, Ossining, USA |
B*39:70 | HG00002937 | JN124207 | Histogenetics, Ossining, USA |
B*39:71 | HG00002938 | JN604854 | Histogenetics, Ossining, USA |
B*39:72 | HG00002942 | JN242432 | Histogenetics, Ossining, USA |
B*39:73 | HG00003107, HG00003108, HG00003109, HG00003110 | JN406452, JN584509, JN584510, JN584512 | Histogenetics, Ossining, USA |
B*40:02:13 | HG00002854 | JN124241 | Histogenetics, Ossining, USA |
B*40:02:14 | HG00002991 | JN205212 | Histogenetics, Ossining, USA |
B*40:02:15 | HG00002992 | JN406411 | Histogenetics, Ossining, USA |
B*40:06:05 | HG00002856 | JN124195 | Histogenetics, Ossining, USA |
B*40:182 | HG00002847 | JN604851 | Histogenetics, Ossining, USA |
B*40:183 | HG00002849, HG00002850 | JN124221, JN124312 | Histogenetics, Ossining, USA |
B*40:184 | HG00002855 | JN182972 | Histogenetics, Ossining, USA |
B*40:185 | HG00002980 | JN418822 | Histogenetics, Ossining, USA |
B*40:186 | HG00002981 | JN124306 | Histogenetics, Ossining, USA |
B*40:186 | HYT | JQ407773 | Oh-Joong Kwon, Seoul, South Korea |
B*40:187 | HG00002982 | JN406454 | Histogenetics, Ossining, USA |
B*40:188 | HG00002987 | JN119423 | Histogenetics, Ossining, USA |
B*40:189 | HG00002989 | JN124275 | Histogenetics, Ossining, USA |
B*40:190 | HG00002990 | JN124313 | Histogenetics, Ossining, USA |
B*40:191 | HG00003116 | JN124262 | Histogenetics, Ossining, USA |
B*40:192 | D252 | JQ316451 | Daniel Fuerst, Ulm, Germany |
B*40:193 | JRC-D01-063 | AB696716 | Kenji Tadokoro, Tokyo, Japan |
B*41:20 | HG00002846 | JN123809 | Histogenetics, Ossining, USA |
B*44:02:23 | HG00002883 | JN124181 | Histogenetics, Ossining, USA |
B*44:02:24 | HG00002885 | JN123801 | Histogenetics, Ossining, USA |
B*44:03:15 | HG00003019 | JN124295 | Histogenetics, Ossining, USA |
B*44:37:02 | HG00003031 | JN584507 | Histogenetics, Ossining, USA |
B*44:144 | HG00002884 | JN182970 | Histogenetics, Ossining, USA |
B*44:145 | HG00002888 | JN584503 | Histogenetics, Ossining, USA |
B*44:146 | HG00003022 | JN124282 | Histogenetics, Ossining, USA |
B*44:147 | HG00003023 | JN124253 | Histogenetics, Ossining, USA |
B*44:148 | HG00003025, HG00003026 | JN119382, JN584508 | Histogenetics, Ossining, USA |
B*44:149N | HG00003032 | JN119369 | Histogenetics, Ossining, USA |
B*44:150 | 2011090109 | JQ413256 | Antonio Balas, Madrid, Spain |
B*45:14 | HG00002986 | JN124256 | Histogenetics, Ossining, USA |
B*50:16 | HG00002851 | JN124187 | Histogenetics, Ossining, USA |
B*51:01:31 | HG00002916 | JN124238 | Histogenetics, Ossining, USA |
B*51:01:32 | HG00002918 | JN584504 | Histogenetics, Ossining, USA |
B*51:01:33 | HG00002921 | JN124219 | Histogenetics, Ossining, USA |
B*51:01:34 | HG00003077 | JN124315 | Histogenetics, Ossining, USA |
B*51:129 | HG00002845 | JN644965 | Histogenetics, Ossining, USA |
B*51:130 | HG00003069 | JN124255 | Histogenetics, Ossining, USA |
B*51:131 | HG00003076 | JN124249 | Histogenetics, Ossining, USA |
B*52:01:11 | HG00003050 | JN205196 | Histogenetics, Ossining, USA |
B*52:28 | HG00002899 | JN124194 | Histogenetics, Ossining, USA |
B*53:01:06 | HG00002891 | JN182973 | Histogenetics, Ossining, USA |
B*53:01:07 | HG00003034 | JN406456 | Histogenetics, Ossining, USA |
B*54:26 | 5301000921 | JN209963 | Chuanfu Zhu, Jinan, China |
B*55:01:09 | HG00002922 | JN124177 | Histogenetics, Ossining, USA |
B*55:56 | HG00003060 | JN124265 | Histogenetics, Ossining, USA |
B*56:34 | HG00003061 | JN124273 | Histogenetics, Ossining, USA |
B*56:35 | JRC-D78-004 | AB696718 | Kenji Tadokoro, Tokyo, Japan |
B*57:56 | HG00003052 | JN561730 | Histogenetics, Ossining, USA |
B*57:57 | MOCEric | HE650819 | Sylvie Tourne, Strasbourg, France |
B*58:01:10 | HG00002900 | JN604852 | Histogenetics, Ossining, USA |
B*58:01:11 | HG00002901 | JN124234 | Histogenetics, Ossining, USA |
C*01:66 | HG00003123 | JN205215 | Histogenetics, Ossining, USA |
C*03:02:09 | HG00003124, HG00003125 | JN124428, JN124445 | Histogenetics, Ossining, USA |
C*03:04:26 | JRC-D01-078 | AB696717 | Kenji Tadokoro, Tokyo, Japan |
C*04:112 | Nov-42 | JQ437409 | Dimitri Monos, Philadelphia, USA |
C*06:74 | HG00003129 | JN124399 | Histogenetics, Ossining, USA |
C*06:75 | HG00003154 | JN561734 | Histogenetics, Ossining, USA |
C*07:02:01:04 | TU05 | AB679917 | Takashi Shiina, Isehara, Japan |
C*07:02:01:05 | TU07 | AB679918 | Takashi Shiina, Isehara, Japan |
C*07:02:30 | HG00003126 | JN629058 | Histogenetics, Ossining, USA |
C*07:228 | HG00003127 | JN124373 | Histogenetics, Ossining, USA |
C*07:229 | HG00003133 | JN124381 | Histogenetics, Ossining, USA |
C*07:230 | HG00003134 | JN584519 | Histogenetics, Ossining, USA |
C*07:231 | HG00003138 | JN124410 | Histogenetics, Ossining, USA |
C*07:232 | HG00003150 | JN561736 | Histogenetics, Ossining, USA |
C*07:233 | HG00003151 | JN124418 | Histogenetics, Ossining, USA |
C*07:234 | HG00003153 | JN205216 | Histogenetics, Ossining, USA |
C*07:235 | FRF2373 | HE653003 | Sylvie Tourne, Strasbourg, France |
C*08:16:02 | HG00003158 | JN406420 | Histogenetics, Ossining, USA |
C*08:60 | HG00003165 | JN124424 | Histogenetics, Ossining, USA |
C*12:71 | HG00003157 | JN124382 | Histogenetics, Ossining, USA |
C*12:72 | HG00003178 | JN629060 | Histogenetics, Ossining, USA |
C*15:02:08 | 90169129 | JQ511980 | Daniel Fuerst, Ulm, Germany |
C*16:45 | HG00003176 | JN124408 | Histogenetics, Ossining, USA |
DRB1*03:80 | 5461000145 | JF896313 | Chuanfu Zhu, Jinan, China |
DRB1*12:16:02 | 2011-1659 | JQ629933 | A-Hyun Lim, Seoul, South Korea |
DRB1*14:54:02 | NT01153, NT01207 | HQ189133, JQ253896 | Carolyn K Hurley, Washington DC, USA |
DRB3*02:29N | LUMC-DRB3N79, 10D05530 | FR823459 | Jacqy DH Anholts, Leiden, The Netherlands |
MICA*070 | USAPET3 | JQ522969 | Effie Petersdorf, Seattle, USA |
MICA*071 | SZGSQ-14 | JQ619159 | Su-Qing Gao, Shenzhen, China |
Table 2: Confirmatory Sequences
Sequence | Cell identification | Accession number | Submitting author |
---|---|---|---|
A*01:37 | DALE740378AN | HE646286 | Steven GE Marsh, London, United Kingdom |
A*02:315 | 55360 | JQ410445 | Jianping Li, Shenyang, Liaoning, China |
B*40:172 | 09-72-6890 | HE650145 | Wendy Swelsen, Amsterdam, The Netherlands |
B*46:30 | 5331000343 | JN209964 | Chuanfu Zhu, Jinan, China |
C*05:55 | Ua2010tp2515 | HE651324 | Mats Bengtsson, Uppsala, Sweden |
DRB1*04:01:02 | 22685461 | HE654651 | Thibaut Gervais, Brussels, Belgium |
DRB1*14:76 | 5993312 | HE603109 | Katherine Poole, Leeds, United Kingdom |
Table 3: Recently Published Sequences
Sequence | References |
---|---|
A*02:330 | (3) |
A*11:108 | (3) |
A*31:53 | (4) |
A*66:14 | (5) |
B*08:01:08 | (6) |
B*15:01:18 | (7) |
B*35:87 | (8) |
B*40:175 | (3) |
B*40:176 | (3) |
B*44:110 | (7) |
B*51:63 | (9) |
C*04:01:22 | (7) |
DQB1*03:03:04 | (10) |
DQB1*05:14 | (7) |
DPB1*06:01 | (11) |
DPB1*10:01 | (11) |
DPB1*11:01:01 | (11) |
DPB1*16:01 | (11) |
DPB1*17:01 | (11) |
DPB1*18:01 | (11) |
DPB1*21:01 | (11) |
DPB1*26:01:02 | (11) |
DPB1*27:01 | (11) |
DPB1*39:01 | (11) |
DPB1*131:01 | (11) |
DPB1*135:01 | (11) |
MICA*010:02 | (12) |
References:
- Marsh SGE, Albert ED, Bodmer WF, et al. Nomenclature for Factors of the HLA system, 2010. Tissue Antigens. 2010 75:291-455.
- Robinson J, Malik A, Parham P, Bodmer JG, Marsh SGE. IMGT/HLA database--a sequence database for the human major histocompatibility complex. Tissue Antigens. 2000; 55: 280-7.
- Endres RO, Hoffman AD, Scott I. Characterization of four novel HLA alleles, including two in the same haplotype. Tissue Antigens. 2012. 79: 204-5.
- Witter K, Kutsch M, Zahn R, Albert T, Dick A, Kauke T. Sequence-based typing confirmed a new HLA-A allele, A*31:53, in a liver recipient patient. Tissue Antigens. 2012. 79: 206-7.
- Wowk PF, Mattar SB, Rodrigues F, da Graca Bicalho M. A novel allele, HLA-A*66:14, identified in a Brazilian volunteer bone marrow donor. Tissue Antigens. 2012. 79: 207-8.
- Schlahsa L, Immenschuh S, Eiz-Vesper B, Blasczyk R, Figueiredo C. HLA-B*08:01:08- joining the fold of silent alpha-1 proline mutations in HLA-B. Tissue Antigens. 2012. 79: 208-9.
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- Buddaseth S, Muller K, Bade-Doeding C, Blasczyk R, Huyton T. The new allele HLA-B*35:87 was generated by recombination between HLA-B*35:01 and B*08:01 in intron 2. Tissue Antigens. 2012. 79: 211-2.
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