Nomenclature for Factors of the HLA System

October 2009

Compiled by Steven G. E. Marsh for the WHO Nomenclature Committee for Factors of the HLA System.

Published In:

The following sequences have been submitted to the Nomenclature Committee since the September 2009 nomenclature update and, following agreed policy, have been assigned official allele designations. Full details of all sequences will be published in a forthcoming report.

Below are listed the newly assigned sequences (Table 1) and confirmations of previously reported sequences (Table 2). The accession number of each sequence is given and these can be used to retrieve the sequence files from the EMBL, GenBank or DDBJ data libraries. Although accession numbers have been assigned by the data-libraries and most sequences are already available, there is still the possibility that an author may not yet have allowed the sequence to be released; in such a case you will have to contact the submitting author directly. Additional information pertaining to new sequences is often included in the publications describing these alleles; a listing of recent publications that describe new HLA sequences is given in Table 3.

All new and confirmatory sequences should now be submitted directly to the WHO Nomenclature Committee for Factors of the HLA System via the IMGT/HLA Database using the sequence submission tool provided. The IMGT/HLA Database may be accessed via the world wide web at : http://ww.ebi.ac.uk/imgt/hla/.

Table 1: New Sequences


Sequence Cell identification Accession number Submitting author
A*010109 HN-98001-3 FJ619449 Histogenetics, Ossining, USA
A*010110 HN-64111-0,
HN-74486-0,
HN-99884-6,
HN-21154-4
FJ619453,
FJ765917,
FJ898483,
FJ976757,
Histogenetics, Ossining, USA
A*010111 HN-1066965 FJ392158 Histogenetics, Ossining, USA
A*010112 HN-74212-0 FJ765916 Histogenetics, Ossining, USA
A*010113 HN-00401-6 FJ875535 Histogenetics, Ossining, USA
A*0146 HN-5357 FJ392149 Histogenetics, Ossining, USA
A*0147 HN-8045382 FJ619460 Histogenetics, Ossining, USA
A*0148 HN-2260722 FJ619470 Histogenetics, Ossining, USA
A*0149 HN-46950-9,
HN-47555-4
FJ765918,
FJ875541,
Histogenetics, Ossining, USA
A*0150 HN-1446866 FJ765935 Histogenetics, Ossining, USA
A*0151 HN-384D FJ785207 Histogenetics, Ossining, USA
A*020133 HN-30259-5 FJ619441 Histogenetics, Ossining, USA
A*020134 HN-79594-0 FJ619451 Histogenetics, Ossining, USA
A*020135 HN-92009-0 FJ619454 Histogenetics, Ossining, USA
A*020136 HN-51291-4 FJ765907 Histogenetics, Ossining, USA
A*020137 HN-66993-7,
HN-60669-8,
HN-58101-4
FJ538268,
FJ549404,
FJ875539,
Histogenetics, Ossining, USA
A*020138 HN-81213-1 FJ549402 Histogenetics, Ossining, USA
A*020139 HN-72740-4 FJ549403 Histogenetics, Ossining, USA
A*020140 HN-673307 FJ594708 Histogenetics, Ossining, USA
A*020141 HN-55832-0 FJ600650 Histogenetics, Ossining, USA
A*020142 HN-86940-0 FJ875534 Histogenetics, Ossining, USA
A*020608 HN-2356550 FJ538269 Histogenetics, Ossining, USA
A*020609 HN-2111765 FJ792529 Histogenetics, Ossining, USA
A*030117 HN-29269-1 FJ619442 Histogenetics, Ossining, USA
A*0363 HN-50847-0,
HN-52450-1
FJ619456,
FJ640590
Histogenetics, Ossining, USA
A*0364 HN-0664550 FJ619462 Histogenetics, Ossining, USA
A*0365 HN-0300840486922 FJ765936 Histogenetics, Ossining, USA
A*0366 HN-47860-9 FJ765940 Histogenetics, Ossining, USA
A*0367 HN-64366-4 FJ875540 Histogenetics, Ossining, USA
A*0368N HN-76532-7 FJ875548 Histogenetics, Ossining, USA
A*0369N HN-89228-9,
HN-89227-1
FJ875549,
FJ875550
Histogenetics, Ossining, USA
A*110115 HN-770624,
HN-2772718,
HN-679700
FJ489877,
FJ765892,
FJ792523
Histogenetics, Ossining, USA
A*110116 HN-98268-4 FJ765938 Histogenetics, Ossining, USA
A*1154 HN-53533-5 FJ358620 Histogenetics, Ossining, USA
A*1155 SZ-38 GU074394 Hong-Yan Zou, Shenzhen, China
A*2323 HN-93402-4 FJ619391 Histogenetics, Ossining, USA
A*2324 HN-61977-5,
HN-02796-4
FJ619400,
FJ619417
Histogenetics, Ossining, USA
A*2325 HN-27733-7 FJ619412 Histogenetics, Ossining, USA
A*240227 HN-8436563,
HN-00138-7
FJ619476,
FJ907388
Histogenetics, Ossining, USA
A*240228 HN-7940633 FJ600640 Histogenetics, Ossining, USA
A*240229 HN-54259-0,
HN-4519810,
HN-29841-4
FJ619411,
FJ619463,
FJ619474
Histogenetics, Ossining, USA
A*240230 HN-4999749,
HN-3333619
FJ619465,
FJ875689
Histogenetics, Ossining, USA
A*2642 HN-8180403 FJ765894 Histogenetics, Ossining, USA
A*2921 HN-68275-4 FJ875544 Histogenetics, Ossining, USA
A*3032 HN-0546437 FJ765886 Histogenetics, Ossining, USA
A*310104 HN-5934489 FJ358622 Histogenetics, Ossining, USA
A*310105 HN-7047696 FJ358632 Histogenetics, Ossining, USA
A*310106 HN-1713861 FJ792524 Histogenetics, Ossining, USA
A*3128 CMHRN1660 FJ785520 Min Yu, Chicago, USA
A*320104 HN-93481-8,
HN-26034-0
FJ619392,
FJ976750
Histogenetics, Ossining, USA
A*3329 HN-7429324 FJ792528 Histogenetics, Ossining, USA
A*6612 HN-19916-4,
HN-07421-8
FJ765944,
FJ785209
Histogenetics, Ossining, USA
A*680203 HN-69877-5 FJ785206 Histogenetics, Ossining, USA
A*680204 HN-710638 FJ792521 Histogenetics, Ossining, USA
A*9296 HN-23582-9 FJ619440 Histogenetics, Ossining, USA
A*9297 HN-12024-6 FJ619447 Histogenetics, Ossining, USA
A*9298 HN-38943-8 FJ619448 Histogenetics, Ossining, USA
A*9299 HN-61873-4 FJ765898 Histogenetics, Ossining, USA
B*080110 HN-0957535,
HN-0957550
FJ765780,
FJ765781
Histogenetics, Ossining, USA
B*0848 HN-77831-6 FJ640581 Histogenetics, Ossining, USA
B*0849 HN-6751512 FJ765784 Histogenetics, Ossining, USA
B*0850 HN-6520188 FJ765768 Histogenetics, Ossining, USA
B*0851 HN-21578-0 FJ765813 Histogenetics, Ossining, USA
B*0852 HN-75383-4 FJ765842 Histogenetics, Ossining, USA
B*0853 HN-31452-7 FJ765854 Histogenetics, Ossining, USA
B*0854 HN-4859149 FJ765868 Histogenetics, Ossining, USA
B*1331 HN-5794422 FJ560457 Histogenetics, Ossining, USA
B*1332 HN-7628141 FJ765550 Histogenetics, Ossining, USA
B*1332 BY00533 GU066745 Carolyn K Hurley, Washington DC, USA
B*1333 HN-3340275 FJ765775 Histogenetics, Ossining, USA
B*1334 HN-0300931018822 FJ858892 Histogenetics, Ossining, USA
B*1335 HN-94049-4 FJ875563 Histogenetics, Ossining, USA
B*140204 HN-3547 FJ765538 Histogenetics, Ossining, USA
B*140205 BY00531 GU066743 Carolyn K Hurley, Washington DC, USA
B*1417 HN-0727001 FJ765955 Histogenetics, Ossining, USA
B*1418 HN-71743-6 FJ875560 Histogenetics, Ossining, USA
B*150113 HN-62698-9 FJ866160 Histogenetics, Ossining, USA
B*150114 HN-8715483,
HN-8715715
FJ875678,
FJ875679
Histogenetics, Ossining, USA
B*150115 HN-9592550,
HN-86512-7
FJ875680,
FJ875552,
FJ875680
Histogenetics, Ossining, USA
B*150116 HN-708D FJ796992 Histogenetics, Ossining, USA
B*150203 HN-3219446 FJ392165 Histogenetics, Ossining, USA
B*150204 HN-675880 FJ468323 Histogenetics, Ossining, USA
B*150502 HN-86737-1 FJ875570 Histogenetics, Ossining, USA
B*151002 HN-6971276 FJ765804 Histogenetics, Ossining, USA
B*151804 HN-00570 FJ765772 Histogenetics, Ossining, USA
B*180106 HN-3233058 FJ494824 Histogenetics, Ossining, USA
B*180107 HN-45151-5 FJ765926 Histogenetics, Ossining, USA
B*180108 HN-7694 FJ875562 Histogenetics, Ossining, USA
B*1846 HN-1713168 FJ765796 Histogenetics, Ossining, USA
B*1847 HN-4276 FJ765819 Histogenetics, Ossining, USA
B*270513 HN-77796-1 FJ765818 Histogenetics, Ossining, USA
B*2760 HN-5161570,
HN-4010737
FJ765791,
FJ765792
Histogenetics, Ossining, USA
B*2761 HN-0826820 FJ765808 Histogenetics, Ossining, USA
B*2762 HN-54802-0 FJ875554 Histogenetics, Ossining, USA
B*350114 HN-5709345 FJ494826 Histogenetics, Ossining, USA
B*350115 HN-1642615,
HN-97678-7
FJ765797,
FJ952582
Histogenetics, Ossining, USA
B*350116 HN-2644283 FJ360522 Histogenetics, Ossining, USA
B*350117 HN-692059 FJ468324 Histogenetics, Ossining, USA
B*350118 HN-7566220 FJ765870 Histogenetics, Ossining, USA
B*350119 HN-18209-4 FJ765985 Histogenetics, Ossining, USA
B*350303 HN-6933607 FJ594718 Histogenetics, Ossining, USA
B*352402 HN-6458538 FJ765558 Histogenetics, Ossining, USA
B*3720 HN-8481 FJ765820 Histogenetics, Ossining, USA
B*3721 HN-66524-7 FJ765972 Histogenetics, Ossining, USA
B*3821 HN-5161976 FJ765541 Histogenetics, Ossining, USA
B*3822 HN-732020 FJ468328 Histogenetics, Ossining, USA
B*390502 HN-3800450 FJ765561 Histogenetics, Ossining, USA
B*3955 HN-1714885 FJ765826 Histogenetics, Ossining, USA
B*3956 HN-7445284 FJ392183 Histogenetics, Ossining, USA
B*400208 HN-90110-6 FJ765982 Histogenetics, Ossining, USA
B*4099 HN-0397427 FJ235025 Histogenetics, Ossining, USA
B*4099 BY00480 GQ410099 Carolyn K Hurley, Washington DC, USA
B*440211 HN-01937-9 FJ765963 Histogenetics, Ossining, USA
B*440212 HN-35487-4 FJ875569 Histogenetics, Ossining, USA
B*440502 HN-78228-0 FJ875553 Histogenetics, Ossining, USA
B*4489 HN-77525-6 FJ866161 Histogenetics, Ossining, USA
B*4490 HN-07064-2 FJ875671 Histogenetics, Ossining, USA
B*4491 HN-4002229 FJ765774 Histogenetics, Ossining, USA
B*4492 HN-48769-2 FJ765839,
FJ765983
Histogenetics, Ossining, USA
B*4493 HN-4857960 FJ765867 Histogenetics, Ossining, USA
B*4494 HN-5715940 FJ765930 Histogenetics, Ossining, USA
B*4495 HN-33336-6 FJ765966 Histogenetics, Ossining, USA
B*4496 HN-33937-0 FJ765975 Histogenetics, Ossining, USA
B*4497 HN-60214-0,
HN-60432-8
FJ765979,
FJ765980
Histogenetics, Ossining, USA
B*4498 HN-74975-2 FJ765981 Histogenetics, Ossining, USA
B*4499 HN-6578988 FJ858898 Histogenetics, Ossining, USA
B*4511 HN-8041258 FJ594700 Histogenetics, Ossining, USA
B*460103 HN-3681660 FJ765777 Histogenetics, Ossining, USA
B*480202 HN-6126954 FJ875672 Histogenetics, Ossining, USA
B*4821 HN-4187485 FJ494825 Histogenetics, Ossining, USA
B*4908 HN-32306-0 FJ765964 Histogenetics, Ossining, USA
B*4909 HN-50261-4 FJ765969 Histogenetics, Ossining, USA
B*510114 HN-1575924 FJ765785 Histogenetics, Ossining, USA
B*510115 HN-5571489 FJ765836 Histogenetics, Ossining, USA
B*510116 HN-41340-7 FJ796990 Histogenetics, Ossining, USA
B*512403 HN-30281-0 FJ765860 Histogenetics, Ossining, USA
B*5185 HN-1736219 FJ765569 Histogenetics, Ossining, USA
B*5186 HN-3623019 FJ765793 Histogenetics, Ossining, USA
B*5187 HN-12571-3 FJ640578 Histogenetics, Ossining, USA
B*5188 HN-44579-8 FJ765967 Histogenetics, Ossining, USA
B*5217 HN-0387022 FJ765823 Histogenetics, Ossining, USA
B*5218 HN-3280919 FJ875668 Histogenetics, Ossining, USA
B*5219 HN-8583366 FJ765767 Histogenetics, Ossining, USA
B*530802 HN-83406-0 FJ765858 Histogenetics, Ossining, USA
B*5320 HN-24211-5 FJ875673 Histogenetics, Ossining, USA
B*5321 BY00536 GU066748 Carolyn K Hurley, Washington DC, USA
B*5539 HN-1576783 FJ765822 Histogenetics, Ossining, USA
B*560103 HN-699D FJ796991 Histogenetics, Ossining, USA
B*5627 HN-51990-6 FJ858895 Histogenetics, Ossining, USA
B*570106 HN-93081-6 FJ866182 Histogenetics, Ossining, USA
B*570202 HN-6130964,
HN-4094699,
HN-711438
FJ502326,
FJ765835,
FJ468327,
Histogenetics, Ossining, USA
B*5727 HN-2155559 FJ765827 Histogenetics, Ossining, USA
B*5728N An188086 FN555101 Francesca Poli, Milano, Italy
B*580104 B10020 GQ452848 Keming Du, Shanghai, China
B*5825 HN-0516510 FJ765786 Histogenetics, Ossining, USA
B*5826 LAAL686132AN FN555526 Steven GE Marsh, London, United Kingdom
B*8203 HN-7535450 FJ765800 Histogenetics, Ossining, USA
B*9581N HN-72542-2,
HN-62535-9
FJ875640,
FJ875647
Histogenetics, Ossining, USA
B*9582N HN-95367-1 FJ875653 Histogenetics, Ossining, USA
B*9583 HN-6117736 FJ765817 Histogenetics, Ossining, USA
B*9584 HN-44736612 FJ765960 Histogenetics, Ossining, USA
B*9585 BY00534 GU066746 Carolyn K Hurley, Washington DC, USA
Cw*0449 328333 FN430611 Tatiana Lebedeva, Dedham, USA
Cw*050111 329141 FN422380 Tatiana Lebedeva, Dedham, USA
Cw*0531 309439 FN422381 Tatiana Lebedeva, Dedham, USA
Cw*0792 356442 FN430613 Tatiana Lebedeva, Dedham, USA
Cw*0793 LUMC-C65 FN555606 Jacqy DH Anholts, Leiden, The Netherlands
Cw*080203 311562 FN422383 Tatiana Lebedeva, Dedham, USA
Cw*0830 347187 FN422384 Tatiana Lebedeva, Dedham, USA
G*010120 BCI-04 GU070582 Jorge Martinez-Laso, Madrid, Spain
DRB1*010116 HN-27144-5 FJ392251 Histogenetics, Ossining, USA
DRB1*010116 BY00546 GU066757 Carolyn K Hurley, Washington DC, USA
DRB1*031102 BY00543 GU066754 Carolyn K Hurley, Washington DC, USA
DRB1*0350 93051421 EU445580 Andrea Vigh, Ulm, Germany
DRB1*0351 BY00538 GU066750 Carolyn K Hurley, Washington DC, USA
DRB1*041702 BY00541 GU066752 Carolyn K Hurley, Washington DC, USA
DRB1*047202 BY00544 GU066755 Carolyn K Hurley, Washington DC, USA
DRB1*0909 BY00548 GU066759 Carolyn K Hurley, Washington DC, USA
DRB1*1185 BY00551 GU066762 Carolyn K Hurley, Washington DC, USA
DRB1*1186 BY00547 GU066758 Carolyn K Hurley, Washington DC, USA
DRB1*1187 BY00539 GU066751 Carolyn K Hurley, Washington DC, USA
DRB1*130106 BY00542 GU066753 Carolyn K Hurley, Washington DC, USA
DRB1*1493 BY00549 GU066760 Carolyn K Hurley, Washington DC, USA
DRB1*1494 BY00540 GU066764 Carolyn K Hurley, Washington DC, USA
DQB1*0404 195212 FN555147 Klaus Witter, München, Germany
DPB1*2702 196660,
197150,
197151
FN555432 Klaus Witter, München, Germany
DPB1*2802 UMKA9360A1 FN555527 Uma Kanga, New Delhi, India

Table 2: Confirmatory Sequences


Sequence Cell identification Accession number Submitting author
A*030114 4VL071332 FN555233 Arturas Jakubauskas, Vilnius, Lithuania
A*9290 HN-83336-8 FJ785211 Histogenetics, Ossining, USA
B*070214 HN-59808-2 FJ765977 Histogenetics, Ossining, USA
B*1833 BY00530 GU066742 Carolyn K Hurley, Washington DC, USA
B*2747 HN-45714-9 FJ796989 Histogenetics, Ossining, USA
B*4109 BY00537 GU066749 Carolyn K Hurley, Washington DC, USA
B*480302 BY00529 GU066741 Carolyn K Hurley, Washington DC, USA
B*5178 AN188087 FN555006 Francesca Poli, Milano, Italy
B*5626 HN-0340062 FJ765799 Histogenetics, Ossining, USA
B*5818 BY00528 GU066740 Carolyn K Hurley, Washington DC, USA
Cw*0127 CMHD296.4 GQ402147 Min Yu, Chicago, USA
Cw*05010102 7550800776 GQ895733 Yunping Xu, Shenzhen, China
Cw*075602 300878 FN422382 Tatiana Lebedeva, Dedham, USA
Cw*0773 333711 FN430612 Tatiana Lebedeva, Dedham, USA
Cw*1523 HN-25224-7 FJ875625 Histogenetics, Ossining, USA
G*01010101 BCI-08,
BCI-09,
BCI-13,
BCI-17,
BCI-18,
CUI-61
GQ996557,
GQ996563
Jorge Martinez-Laso, Madrid, Spain
G*01010105 BCI-10,
CUI-61,
CUI-80
GQ996565 Jorge Martinez-Laso, Madrid, Spain
G*01010201 BCI-13 GQ996569 Jorge Martinez-Laso, Madrid, Spain
G*010108 CUI-62 GQ996568 Jorge Martinez-Laso, Madrid, Spain
G*0103 BCI-06,
BCI-08
GQ996561,
GQ996562
Jorge Martinez-Laso, Madrid, Spain
G*010401 BCI-19 GQ996559 Jorge Martinez-Laso, Madrid, Spain
G*010403 CUI-62 GQ996567 Jorge Martinez-Laso, Madrid, Spain
G*010404 BCI-18 GQ996558 Jorge Martinez-Laso, Madrid, Spain
G*0106 BCI-09 GQ996564 Jorge Martinez-Laso, Madrid, Spain
DRB1*0479 BY00545 GU066756 Carolyn K Hurley, Washington DC, USA
DRB1*0707 41618 FN556019 Thibaut Gervais, Brussels, Belgium
DRB1*1169 BY00550 GU066761 Carolyn K Hurley, Washington DC, USA
DRB1*1172 BY00527 GU066739 Carolyn K Hurley, Washington DC, USA
DRB1*1615 BY00552 GU066763 Carolyn K Hurley, Washington DC, USA
DRB3*0220 63140 FN556018 Jacqy DH Anholts, Leiden, The Netherlands
DRB3*0303 48280,
48733
FN556017 Jacqy DH Anholts, Leiden, The Netherlands

Table 3: Recently Published Sequences


Sequence References
A*0260 (1)
A*030106 (2)
A*030108 (3)
A*0337 (2)
A*110203 (4)
A*2317 (2)
A*2480 (2)
A*2491 (3)
A*2498 (3)
A*3023 (2)
A*330303 (3)
A*3317 (3)
A*680202 (1)
A*9225N (2)
A*9234 (2)
B*070206 (2)
B*0759 (2)
B*0761 (2)
B*0765 (2)
B*132202 (5)
B*150106 (2)
B*1827 (2)
B*352002 (2)
B*3585 (2)
B*3818 (6)
B*3943 (2)
B*4096 (7)
B*5151 (2)
B*5401 (8)
B*5417 (8)
B*5419 (9)
B*4082 (2)
B*9521 (10)
B*9549N (11)
Cw*0121 (12)
Cw*030301 (13)
Cw*0342 (2)
Cw*034301 (2)
Cw*0344 (2)
Cw*0428 (2)
Cw*04010103 (14)
Cw*0430 (2)
Cw*0433 (2)
Cw*050104 (2)
Cw*0519 (2)
Cw*060203 (2)
Cw*070109 (2)
Cw*070202 (2)
Cw*070205 (15)
Cw*070206 (16)
Cw*0750 (2)
Cw*0815 (2)
Cw*0827 (17)
Cw*120306 (2)
Cw*1409 (2)
G*01010301 (18)
G*01010302 (18)
DRB1*0336 (2)
DRB1*0473 (2)
DRB1*0717 (19)
DRB1*120204 (20)
DRB1*1382 (2)

References:

  1. Serov Y, Hommel-Berrey G, Lobashevsky A. Identification of a novel HLA-A*680202 allele by sequence-based typing in an African American individual. Tissue Antigens. 2009 74:435-7.
  2. Anholts JD, Aneq M, Dirks HL, Tas A, Verduyn W, Oudshoorn M. Thirty-six novel HLA alleles: 7 HLA-A, 11 HLA-B, 15 HLA-C and 3 HLA-DRB1. Tissue Antigens. 2009 74:424-8.
  3. Yan LX, Zhu FM, He J, Zhang W. Five novel HLA-A alleles, HLA-A*030108, A*2491, A*2498, A*330303, A*3317 were identified by polymerase chain reaction sequence based typing. Tissue Antigens. 2009 74:432-4.
  4. Li Z, Zou HY, Wang DM, Cheng X. A novel allele, HLA-A*110203, was identified in a Chinese individual. Tissue Antigens. 2009 74:434-5.
  5. Zou HY, Cheng LH, Li Z, Jin SZ. Full length sequence of a novel HLA-B*132202 allele. Tissue Antigens. 2009 74:437-9.
  6. Cheng LH, Zhu WG, Lan YX, Jin SZ, Zou HY. Full length sequence of a novel HLA-B*3818 allele differs from HLA-B*380201 allele in exon 4 and intron 5. Tissue Antigens. 2009 74:439-40.
  7. Zhang Y, Lu HX, Liu Y, Zhu CF. A novel HLA-B allele, B*4096 was identified by sequence-based typing in a Chinese individual. Tissue Antigens. 2009 74:440-2.
  8. Cheng LH, Zhu WG, Lan YX, Zou HY, Li Z, Jin SZ. A novel HLA-B*54 allele, B*5417, identified in a Northern Chinese Han individual. Tissue Antigens. 2009 74:442-4.
  9. Lee KW, Seo JJ. A novel HLA-B allele: HLA-B*5419. Tissue Antigens. 2009 74:444-5.
  10. Chu CC, Chen YF, Lee HL, Lin CY, Lin M. Identification of HLA-B*9521 in Taiwan. Tissue Antigens. 2009 74:445-6.
  11. Zou HY, Li Z, Jin SZ, Cheng X, Cheng LH. A frame shift due to a two-nucleotide deletion results in an HLA-B null allele, B*9549N. Tissue Antigens. 2009 74:447-9.
  12. Li Z, Zhang H, Jin SZ. Identification of a new HLA-Cw*01 variant, Cw*0121, by sequence-based typing in a Chinese patient. Tissue Antigens. 2009 74:449-51.
  13. Xu Y, Deng Z, Zeng J, Gao S, Yang B. Genomic full length sequence of HLA-Cw*030301, identified by cloning and sequencing. Tissue Antigens. 2009 74:452-3.
  14. Xu Y, Deng Z, Zeng J, Gao S, Wang D. A novel HLA-Cw*04 allele, Cw*04010103, identified by genomic full length cloning and sequencing. Tissue Antigens. 2009 74:453-5.
  15. Zou HY, Jin SZ, Li Z, Cheng LH. Full-length sequence of a novel allele, HLA-Cw*070205. Tissue Antigens. 2009 74:455-6.
  16. Wang DM, Gao SQ, Xu YP, Deng ZH. Identification of a novel HLA-Cw*070206 allele. Tissue Antigens. 2009 74:457-8.
  17. Deng ZH, Wang DM, Xu YP. Cloning and sequencing analysis of a novel HLA-Cw*08 variant allele, Cw*0827. Tissue Antigens. 2009 74:458-60.
  18. Cervera I, Herraiz MA, Roman A, Vidart J, Martinez-Laso J. The novel HLA-G*01010302 allele differs from G*01010301 by a single nucleotide change in intron 5. Tissue Antigens. 2009 74:463-4.
  19. Wurm M, Tischer S, Immenschuh S, Blasczyk R, Eiz-Vesper B. The predicted protein structures of the novel DRB1*0717 and DRB1*0701 are highly related. Tissue Antigens. 2009 74:460-2.
  20. Ji Y, Sun Y, Xie JH, Yang JH, Liu Y, Du KM. A novel HLA-DRB1*120204 allele found in a Chinese individual. Tissue Antigens. 2009 74:462-3.