DRB4 References
Assigned October 2024
| Allele Name | Cell ID | HLA ID | Authors | Title | Journal | Year | Vol. | Pages | Contact |
|---|---|---|---|---|---|---|---|---|---|
| DRB4*01:01:01:01 | FH05 | HLA00905 | Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE | Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. | HLA | 2022 | 99 | 328-356 | Prof. Steven GE Marsh |
| DRB4*01:01:01:01 | BURKHARDT | HLA00905 | Gregersen PK, Moriuchi T, Karr RW, Obata F, Moriuchi J, Maccari J, Goldberg D, Winchester RJ, Silver J | Polymorphism of HLA-DR beta chains in DR4, -7, and -9 haplotypes: implications for the mechanisms of allelic variation | Proceedings of the National Academy of Sciences USA | 1986 | 83 | 9149-9153 | Dr. Peter Gregersen |
| DRB4*01:01:01:01 | FS LKT3 |
HLA00905 | Gregersen PK, Shen M, Song QL, Merryman P, Degar S, Seki T, Maccari J, Goldberg D, Murphy H, Schwenzer J, Wang CY, Winchester RJ, Nepom GT, Silver J | Molecular diversity of HLA-DR4 haplotypes | Proceedings of the National Academy of Sciences USA | 1986 | 83 | 2642-2646 | Dr. Peter Gregersen |
| DRB4*01:01:01:01 | MOLT-4 | HLA00905 | Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE | Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. | HLA | 2022 | 99 | 328-356 | Prof. Steven GE Marsh |
| DRB4*01:01:01:01 | PRIESS | HLA00905 | Spies T, Sorrentino R, Boss JM, Okada K, Strominger JL | Structural organization of the DR subregion of the human major histocompatibility complex | Proceedings of the National Academy of Sciences USA | 1985 | 82 | 5165-5169 | Prof. Jack L Strominger |
| DRB4*01:01:01:01 | MOU | HLA00905 | Young JA, Wilkinson D, Bodmer WF, Trowsdale J | Sequence and evolution of HLA-DR7- and -DRw53-associated beta-chain genes | Proceedings of the National Academy of Sciences USA | 1987 | 84 | 4929-4933 | Prof. John Trowsdale |
| DRB4*01:01:01:01 | MOU | HLA00905 | Horton R, Gibson R, Coggill P, Miretti M, Allcok RJ, Almeida J, Forbes S, Gilbert JGR, Halls K, Harrow JL, Hart E, Howe K, Jackson DK, Palmer S, Roberts AN, Sims S, Stewart CA, Traherne JA, Trevanion S, Wilming L, Rogers J, de Jong PJ, Elliott JF, Sawcer S, Todd JA, Trowsdale J, Beck S | Variation analysis and gene annotation of eight MHC haplotypes: The MHC Haplotype Project. | Immunogenetics | 2008 | 60 | 1-18 | Prof. Stephan Beck |
| DRB4*01:01:01:01 | MMCC | HLA00905 | Horn GT, Bugawan TL, Long CM, Manos MM, Erlich HA | Sequence analysis of HLA class II genes from insulin-dependent diabetic individuals | Human Immunology | 1988 | 21 | 249-263 | Dr. Henry Erlich |
| DRB4*01:01:01:01 | DM24 DM29 |
HLA00905 | Todd JA, Bell JI, McDevitt HO | HLA-DQ beta gene contributes to susceptibility and resistance to insulin-dependent diabetes mellitus | Nature | 1987 | 329 | 599-604 | Dr. John Todd |
| DRB4*01:01:01:03 | ARBO | HLA23372 | Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE | Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. | HLA | 2022 | 99 | 328-356 | Prof. Steven GE Marsh |
| DRB4*01:01:01:04 | SHJO | HLA31732 | Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE | Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. | HLA | 2022 | 99 | 328-356 | Prof. Steven GE Marsh |
| DRB4*01:01:02 | HG00012473 | HLA15376 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:01:02 | HG00011755 | HLA15376 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:01:03 | HG00018101 | HLA22240 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:01:03 | HG00022266 | HLA22240 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:01:03 | HG00011750 HG00011749 |
HLA22240 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:01:03 | HG00031348 HG00017242 HG00013822 HG00022264 HG00022265 |
HLA22240 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:01:04 | HG00018613 | HLA22373 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:01:04 | HG00025376 | HLA22373 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:01:04 | c2128591 | HLA22373 | - | - | - | - | - | - | Dr. Francisco de Paula Sanchez Gordo |
| DRB4*01:01:04 | PN39817732AN | HLA22373 | - | - | - | - | - | - | Prof. Steven GE Marsh |
| DRB4*01:01:04 | HG00011286 | HLA22373 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:01:04 | HG00014681 HG00018618 HG00018103 HG00015913 HG00022291 HG00022275 HG00015914 HG00016608 HG00014619 HG00014683 |
HLA22373 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:01:04 | HG00025623 HG00025624 |
HLA22373 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:01:04 | HG00022292 | HLA22373 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:01:05 | HG00018611 | HLA22344 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:01:05 | HG00014618 | HLA22344 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:01:06 | 81196967 | HLA22452 | - | - | - | - | - | - | Dr. Daniel Fuerst |
| DRB4*01:01:07 | HG00012949 | HLA23515 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:01:07 | HG00019609 | HLA23515 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:01:07 | HG00022267 | HLA23515 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:01:08 | HG00022286 | HLA23595 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:01:09 | HG00027341 | HLA25824 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:01:09 | HG00022561 | HLA25824 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:01:10 | HG00029058 | HLA30109 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:01:10 | HG00031774 | HLA30109 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:01:11 | 23032847726HS2680 | HLA37518 | - | - | - | - | - | - | Dr. Cathi Murphey |
| DRB4*01:01:11 | HS2680 | HLA37518 | - | - | - | - | - | - | Dr. Cathi Murphey |
| DRB4*01:01:12 | 2210491407 | HLA37807 | Cargou M, Andreani M, Lucas JAM, Wojciechowski E, Visentin J | Characterisation of the novel HLA-DRB4*01:01:12 allele by sequencing-based typing. | HLA | 2024 | 103 | - | Dr. Jonathan Visentin |
| DRB4*01:01:13 | P14847-1 | HLA39861 | - | - | - | - | - | - | Prof. Faming Zhu |
| DRB4*01:02 | C.M.L. | HLA00907 | Buyse I, Emonds MP, Bouillon R, Marynen P, Cassiman JJ | Novel class II HLA-DRB4 and DPB1 alleles found in the Belgian population | Immunogenetics | 1993 | 38 | 380- | Prof. Jean-Jacques Cassiman |
| DRB4*01:02 | C.M.L. | HLA00907 | Naruse TK, Ando R, Nose Y, Kagiya M, Ando H, Kawata H, Nabeya N, Isshiki G, Inoko H | HLA-DRB4 genotyping by PCR-RFLP: diversity in the associations between HLA-DRB4 and DRB1 alleles | Tissue Antigens | 1997 | 49 | 152-159 | Prof. Hidetoshi Inoko |
| DRB4*01:02 | C.M.L. | HLA00907 | Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE | Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. | HLA | 2022 | 99 | 328-356 | Prof. Steven GE Marsh |
| DRB4*01:03:01:01 | HSF7 | HLA00908 | - | - | - | - | - | - | Prof. Stephan Beck |
| DRB4*01:03:01:01 | G081 | HLA00908 | De Pablo R, Solis R, Balas A, Vilches C | Specific amplification of the HLA-DRB4 gene from c-DNA. Complete coding sequence of the HLA alleles DRB4*0103101 and DRB4*01033 | Tissue Antigens | 2002 | 59 | 44-46 | Dr. Rosario de Pablo |
| DRB4*01:03:01:01 | MJ4 | HLA00908 | Curtsinger JM, Hilden JM, Cairns JS, Bach FH | Evolutionary and genetic implications of sequence variation in two nonallelic HLA-DR beta-chain cDNA sequences | Proceedings of the National Academy of Sciences USA | 1987 | 84 | 209-213 | Prof. Fritz H Bach |
| DRB4*01:03:01:01 | MLF | HLA00908 | Horton R, Gibson R, Coggill P, Miretti M, Allcok RJ, Almeida J, Forbes S, Gilbert JGR, Halls K, Harrow JL, Hart E, Howe K, Jackson DK, Palmer S, Roberts AN, Sims S, Stewart CA, Traherne JA, Trevanion S, Wilming L, Rogers J, de Jong PJ, Elliott JF, Sawcer S, Todd JA, Trowsdale J, Beck S | Variation analysis and gene annotation of eight MHC haplotypes: The MHC Haplotype Project. | Immunogenetics | 2008 | 60 | 1-18 | Prof. Stephan Beck |
| DRB4*01:03:01:02N | DBB | HLA00909 | Sutton VR, Knowles RW | An aberrant DRB4 null gene transcript is found that could encode a novel HLA-DR beta chain | Immunogenetics | 1990 | 31 | 112-117 | Dr. Robert W Knowles |
| DRB4*01:03:01:02N | DBB | HLA00909 | Naruse TK, Ando R, Nose Y, Kagiya M, Ando H, Kawata H, Nabeya N, Isshiki G, Inoko H | HLA-DRB4 genotyping by PCR-RFLP: diversity in the associations between HLA-DRB4 and DRB1 alleles | Tissue Antigens | 1997 | 49 | 152-159 | Prof. Hidetoshi Inoko |
| DRB4*01:03:01:02N | DBB | HLA00909 | Sutton VR, Knowles RW | An aberrant DRB4 null gene transcript is found that could encode a novel HLA-DR beta chain | Immunogenetics | 1990 | 31 | 112-117 | Dr. Robert W Knowles |
| DRB4*01:03:01:02N | DBB | HLA00909 | - | - | - | - | - | - | Prof. Carolyn K Hurley |
| DRB4*01:03:01:02N | DBB | HLA00909 | Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE | Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. | HLA | 2022 | 99 | 328-356 | Prof. Steven GE Marsh |
| DRB4*01:03:01:02N | WIN | HLA00909 | Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE | Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. | HLA | 2022 | 99 | 328-356 | Prof. Steven GE Marsh |
| DRB4*01:03:01:02N | MYE 2004 | HLA00909 | - | - | - | - | - | - | Mr. Konstantinos Barsakis |
| DRB4*01:03:01:02N | MYE 2004 | HLA00909 | Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE | Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. | HLA | 2022 | 99 | 328-356 | Prof. Steven GE Marsh |
| DRB4*01:03:01:02N | BY01567 BY01568 |
HLA00909 | - | - | - | - | - | - | Prof. Carolyn K Hurley |
| DRB4*01:03:01:03 | SSTO | HLA06603 | Horton R, Gibson R, Coggill P, Miretti M, Allcok RJ, Almeida J, Forbes S, Gilbert JGR, Halls K, Harrow JL, Hart E, Howe K, Jackson DK, Palmer S, Roberts AN, Sims S, Stewart CA, Traherne JA, Trevanion S, Wilming L, Rogers J, de Jong PJ, Elliott JF, Sawcer S, Todd JA, Trowsdale J, Beck S | Variation analysis and gene annotation of eight MHC haplotypes: The MHC Haplotype Project. | Immunogenetics | 2008 | 60 | 1-18 | Prof. Stephan Beck |
| DRB4*01:03:01:03 | SSTO | HLA06603 | Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE | Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. | HLA | 2022 | 99 | 328-356 | Prof. Steven GE Marsh |
| DRB4*01:03:01:03 | FH12 | HLA06603 | Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE | Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. | HLA | 2022 | 99 | 328-356 | Prof. Steven GE Marsh |
| DRB4*01:03:01:04 | BH | HLA20773 | Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE | Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. | HLA | 2022 | 99 | 328-356 | Prof. Steven GE Marsh |
| DRB4*01:03:01:04 | LBUF LBF |
HLA20773 | Bell JI, Denney D Jr, Foster L, Belt T, Todd JA, McDevitt HO | Allelic variation in the DR subregion of the human major histocompatibility complex | Proceedings of the National Academy of Sciences USA | 1987 | 84 | 6234-6248 | Prof. John Bell |
| DRB4*01:03:01:04 | LBUF LBF |
HLA20773 | - | - | - | - | - | - | Prof. Steven GE Marsh |
| DRB4*01:03:01:04 | SHJO | HLA20773 | Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE | Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. | HLA | 2022 | 99 | 328-356 | Prof. Steven GE Marsh |
| DRB4*01:03:01:04 | EAV, AC | HLA20773 | Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE | Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. | HLA | 2022 | 99 | 328-356 | Prof. Steven GE Marsh |
| DRB4*01:03:01:05 | KY | HLA20628 | Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE | Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. | HLA | 2022 | 99 | 328-356 | Prof. Steven GE Marsh |
| DRB4*01:03:01:05 | LKT3 | HLA20628 | Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE | Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. | HLA | 2022 | 99 | 328-356 | Prof. Steven GE Marsh |
| DRB4*01:03:01:05 | PSCDA0442 | HLA20628 | - | - | - | - | - | - | Dr. Seik-Soon Khor |
| DRB4*01:03:01:06 | SYSU005 | HLA21590 | - | - | - | - | - | - | Dr. Riga Wu |
| DRB4*01:03:01:07 | CHI 007 | HLA21306 | Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE | Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. | HLA | 2022 | 99 | 328-356 | Prof. Steven GE Marsh |
| DRB4*01:03:01:08 | T7527 | HLA21599 | Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE | Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. | HLA | 2022 | 99 | 328-356 | Prof. Steven GE Marsh |
| DRB4*01:03:01:08 | PSCDA0423 | HLA21599 | - | - | - | - | - | - | Dr. Seik-Soon Khor |
| DRB4*01:03:01:09 | DEU | HLA22658 | Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE | Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. | HLA | 2022 | 99 | 328-356 | Prof. Steven GE Marsh |
| DRB4*01:03:01:10 | YAR | HLA22663 | Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE | Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. | HLA | 2022 | 99 | 328-356 | Prof. Steven GE Marsh |
| DRB4*01:03:01:11 | BM14 | HLA22666 | Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE | Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. | HLA | 2022 | 99 | 328-356 | Prof. Steven GE Marsh |
| DRB4*01:03:01:11 | BOLETH | HLA22666 | Andersson G, Larhammar D, Widmark E, Servenius B, Peterson PA, Rask L | Class II genes of the human major histocompatibility complex. Organization and evolutionary relationship of the DR beta genes | Journal of Biological Chemistry | 1987 | 262 | 8748-8758 | Dr. Goran Andersson |
| DRB4*01:03:01:11 | BOLETH | HLA22666 | - | - | - | - | - | - | Prof. Steven GE Marsh |
| DRB4*01:03:01:11 | JHAF | HLA22666 | Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE | Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. | HLA | 2022 | 99 | 328-356 | Prof. Steven GE Marsh |
| DRB4*01:03:01:12 | BUR, E | HLA23081 | Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE | Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. | HLA | 2022 | 99 | 328-356 | Prof. Steven GE Marsh |
| DRB4*01:03:01:13N | HB19 | HLA26755 | - | - | - | - | - | - | Dr. Riga Wu |
| DRB4*01:03:01:14 | MANIKA | HLA29115 | Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE | Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. | HLA | 2022 | 99 | 328-356 | Prof. Steven GE Marsh |
| DRB4*01:03:01:15 | FB11 | HLA30796 | Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE | Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. | HLA | 2022 | 99 | 328-356 | Prof. Steven GE Marsh |
| DRB4*01:03:01:16 | BH | HLA31733 | Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE | Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. | HLA | 2022 | 99 | 328-356 | Prof. Steven GE Marsh |
| DRB4*01:03:01:17 | EC | HLA31515 | Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE | Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. | HLA | 2022 | 99 | 328-356 | Prof. Steven GE Marsh |
| DRB4*01:03:01:18 | AN302594 | HLA38110 | - | - | - | - | - | - | Prof. Steven GE Marsh |
| DRB4*01:03:01:19 | PSCDA1028 | HLA41432 | - | - | - | - | - | - | Dr. Seik-Soon Khor |
| DRB4*01:03:01:20 | PSCDA1146 | HLA41376 | - | - | - | - | - | - | Dr. Seik-Soon Khor |
| DRB4*01:03:02:01 | LKT14 | HLA00910 | Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE | Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. | HLA | 2022 | 99 | 328-356 | Prof. Steven GE Marsh |
| DRB4*01:03:02:01 | W778R | HLA00910 | - | - | - | - | - | - | Dr. Lee Ann Baxter-Lowe |
| DRB4*01:03:02:01 | DKB | HLA00910 | Bell JI, Denney D Jr, Foster L, Belt T, Todd JA, McDevitt HO | Allelic variation in the DR subregion of the human major histocompatibility complex | Proceedings of the National Academy of Sciences USA | 1987 | 84 | 6234-6248 | Prof. John Bell |
| DRB4*01:03:02:01 | DKB | HLA00910 | Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE | Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. | HLA | 2022 | 99 | 328-356 | Prof. Steven GE Marsh |
| DRB4*01:03:02:01 | FKD4010302E1 | HLA00910 | - | - | - | - | - | - | Mr. Konstantinos Barsakis |
| DRB4*01:03:02:01 | PSCDA1100 | HLA00910 | - | - | - | - | - | - | Dr. Seik-Soon Khor |
| DRB4*01:03:02:02 | GRC-212 | HLA31794 | Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE | Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. | HLA | 2022 | 99 | 328-356 | Prof. Steven GE Marsh |
| DRB4*01:03:03 | MUDGH | HLA01225 | - | - | - | - | - | - | Prof. Gottfried Fischer |
| DRB4*01:03:03 | MG-CV | HLA01225 | Voorter CE, Hepkema BG, Lems SP, van den Berg-Loonen EM | Identification of three new DRB1 alleles, DRB1*0107, *0425 and *13012 and confirmation of DRB4*01033 | Tissue Antigens | 2003 | 61 | 398-402 | Prof. Ella M. Van den Berg-Loonen |
| DRB4*01:03:03 | FOA2362 | HLA01225 | Palou E, Nogues N, Gil J, Ribera A | Identification of a new HLA-DRB4 allele (DRB4*01033) by PCR-SSP and direct sequencing | Tissue Antigens | 2000 | 56 | 279-281 | Dr. Eduard Palou |
| DRB4*01:03:03 | G081 | HLA01225 | De Pablo R, Solis R, Balas A, Vilches C | Specific amplification of the HLA-DRB4 gene from c-DNA. Complete coding sequence of the HLA alleles DRB4*0103101 and DRB4*01033 | Tissue Antigens | 2002 | 59 | 44-46 | Dr. Rosario de Pablo |
| DRB4*01:03:04 | 14242 | HLA01594 | Fasano ME, Dametto E, Day S, Dunn P, Tacconella M, Curtoni ES | A new allele, HLA-DRB4*010304 | Tissue Antigens | 2003 | 62 | 76-78 | Dr. Maria Edvige Fasano |
| DRB4*01:03:05 | 113757 | HLA17334 | - | - | - | - | - | - | Dr. Ute Geigenmuller |
| DRB4*01:03:05 | HG00012723 | HLA17334 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:05 | HG00031350 | HLA17334 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:06 | 81172337 | HLA18069 | - | - | - | - | - | - | Dr. Daniel Fuerst |
| DRB4*01:03:07 | HG00011760 | HLA19800 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:07 | HG00017244 | HLA19800 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:07 | HG00031349 | HLA19800 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:07 | HG00017502 HG00017504 |
HLA19800 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:07 | HG00025630 | HLA19800 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:07 | HG00013831 | HLA19800 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:07 | HG00017508 HG00017503 |
HLA19800 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:07 | HG00018096 | HLA19800 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:07 | HG00022281 HG00022262 |
HLA19800 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:07 | HG00022279 | HLA19800 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:08 | HG00017506 | HLA20967 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:08 | HG00031355 | HLA20967 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:09 | HG00019054 | HLA20759 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:09 | HG00013481 | HLA20759 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:09 | HG00017507 | HLA20759 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:09 | HG00025622 | HLA20759 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:09 | HG00015915 HG00016901 HG00017235 HG00018851 |
HLA20759 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:09 | HG00022261 | HLA20759 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:09 | HG00022280 | HLA20759 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:09 | HG00022295 | HLA20759 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:09 | HG00022299 | HLA20759 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:10 | HG00018102 | HLA21092 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:11 | HG00018852 | HLA21543 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:11 | HG00012201 | HLA21543 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:11 | HG00022268 | HLA21543 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:12 | HG00013483 | HLA23180 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:12 | HG00022297 | HLA23180 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:13 | HG00017510 | HLA23123 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:13 | HG00018614 HG00018616 HG00018853 HG00018854 HG00018855 HG00018857 |
HLA23123 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:13 | HG00018615 | HLA23123 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:13 | HG00025375 HG00025629 HG00025627 |
HLA23123 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:13 | HG00025625 | HLA23123 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:13 | HG00025628 | HLA23123 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:13 | HG00015296 HG00013824 HG00016902 |
HLA23123 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:13 | HG00018856 | HLA23123 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:14 | HG00013832 | HLA23087 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:14 | HG00017512 | HLA23087 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:15 | HG00022277 | HLA24064 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:16 | HG00027354 | HLA24498 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:16 | HG00011770 | HLA24498 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:16 | HG00024988 | HLA24498 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:17 | HG00029085 | HLA25538 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:17 | HG00025061 | HLA25538 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:17 | HG00025852 | HLA25538 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:18 | HG00025631 | HLA25916 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:19 | HG00026852 | HLA28544 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:20 | GSQ-58 | HLA29793 | - | - | - | - | - | - | Prof. Su-Qing Gao |
| DRB4*01:03:20 | 21-125-029434 | HLA29793 | - | - | - | - | - | - | Dr. Qian Hu |
| DRB4*01:03:21 | HG00029063 | HLA30481 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:22 | HG00030015 | HLA31560 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:23 | TUJM336 | HLA31941 | - | - | - | - | - | - | Prof. Takashi Shiina |
| DRB4*01:03:23 | HG00017236 | HLA31941 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:24 | HG00030145 | HLA31976 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:25 | HG00030144 | HLA31991 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:26 | HG00022288 | HLA34369 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:26 | HG00011289 | HLA34369 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:26 | HG00013938 | HLA34369 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:26 | HG00031981 | HLA34369 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:27 | HG00015695 | HLA34900 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:27 | HG00031982 | HLA34900 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:28 | HG00050633 | HLA36615 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:29 | HG00050476 | HLA36797 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:30 | 922900123 | HLA36817 | - | - | - | - | - | - | Dr. Dolores Planelles |
| DRB4*01:03:31 | HG00050768 | HLA36889 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:32 | HG00050859 HG00050860 |
HLA37099 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:33 | 23032147481HS26773 | HLA37511 | - | - | - | - | - | - | Dr. Cathi Murphey |
| DRB4*01:03:33 | HS26773HK | HLA37511 | - | - | - | - | - | - | Dr. Cathi Murphey |
| DRB4*01:03:34 | HG00051164 | HLA37796 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:03:35 | 20029650 | HLA38781 | - | - | - | - | - | - | Dr. Silvia Giuliodori |
| DRB4*01:03:35 | AG2429 | HLA38781 | - | - | - | - | - | - | Dr. Aiwen Zhang |
| DRB4*01:03:36 | 404016891 | HLA40227 | - | - | - | - | - | - | Ms. Gabriella Camerini Maciel |
| DRB4*01:04 | 69-218 76-394 |
HLA00911 | De Canck I, Demanet C, Mersch G, Jannes G, Rombout A, Rossau R, Van Waeyenberge C | Characterization of a new DRB4 allele (DRB4*0104) | Tissue Antigens | 1996 | 48 | 213-216 | Dr. Guy Mersch |
| DRB4*01:04 | TBC-4975 | HLA00911 | - | - | - | - | - | - | Dr. Masahiro Satake |
| DRB4*01:05 | 17345 | HLA00912 | Voorter CE, Emonds MP, van den Berg-Loonen EM | Identification of a new DRB4 allele (DRB4*0105) by sequence-based typing | Tissue Antigens | 1997 | 49 | 662-664 | Prof. Ella M. Van den Berg-Loonen |
| DRB4*01:06 | MKOST | HLA01559 | - | - | - | - | - | - | Dr. Heather Dunckley |
| DRB4*01:06 | 116371 | HLA01559 | - | - | - | - | - | - | Dr. Francisco de Paula Sanchez Gordo |
| DRB4*01:07:01 | X0002652 X0002630 X0002601 |
HLA01891 | - | - | - | - | - | - | Dr. Lia Mele |
| DRB4*01:07:02 | 122192 | HLA15019 | - | - | - | - | - | - | Dr. Ute Geigenmuller |
| DRB4*01:07:02 | HG00011763 | HLA15019 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:07:02 | HG00013046 HG00013045 |
HLA15019 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:08 | Tokushima | HLA04572 | Lee E, Ozaki S, Ishii H, Takimoto T, Matsumoto T | A novel allele, HLA-DRB1*01:08, identified by sequence-based typing | Tissue Antigens | 2011 | 78 | 158-159 | Ms. Etsukp Lee |
| DRB4*01:09 | LUMC-DRB4-100 | HLA13511 | - | - | - | - | - | - | Miss. Jacqy DH Anholts |
| DRB4*01:10 | LUMC-DRB4-114 | HLA14114 | - | - | - | - | - | - | Miss. Jacqy DH Anholts |
| DRB4*01:11 | HG00011972 | HLA14659 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:11 | HG00011278 | HLA14659 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:11 | 109551 | HLA14659 | - | - | - | - | - | - | Dr. Ute Geigenmuller |
| DRB4*01:12 | HG00011280 | HLA14660 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:13 | HG00011973 | HLA14661 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:13 | HG00011282 | HLA14661 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:13 | HG00012474 | HLA14661 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:14N | HG00011979 HG00011980 HG00011976 HG00011977 HG00011974 HG00011983 HG00011981 HG00011975 HG00011982 HG00011978 |
HLA14662 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:14N | 93053764 | HLA14662 | - | - | - | - | - | - | Dr. Daniel Fuerst |
| DRB4*01:14N | HG00011285 HG00011284 HG00011283 |
HLA14662 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:14N | 19205151 | HLA14662 | Fuerst D, Tsamadou C, Gowdavally S, Schrezenmeier H, Mytilineos J | HLA-DRB4*01:14 is a null allele, and renamed HLA-DRB4*01:14N | HLA | 2020 | 95 | 73-75 | Dr. Daniel Fuerst |
| DRB4*01:15 | HG00011287 | HLA14663 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:16N | HG00014706 HG00014705 |
HLA14664 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:16N | HG00011288 | HLA14664 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:16N | 076099 | HLA14664 | - | - | - | - | - | - | Dr. Ute Geigenmuller |
| DRB4*01:16N | HG00013048 | HLA14664 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:17 | HG00011984 HG00011992 HG00011995 HG00011991 HG00011985 HG00011990 HG00011989 HG00011986 HG00011987 HG00011994 HG00011996 HG00011993 HG00011988 |
HLA14665 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:17 | HG00015815 HG00015814 |
HLA14665 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:17 | HG00011290 HG00011291 |
HLA14665 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:17 | HG00012487 HG00012486 |
HLA14665 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:17 | HG00013060 | HLA14665 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:18 | HG00011500 | HLA15089 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:19 | HG00011501 | HLA15090 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:20 | HG00011502 | HLA15091 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:21 | HG00011503 | HLA15092 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:22 | 046670 | HLA15015 | - | - | - | - | - | - | Dr. Ute Geigenmuller |
| DRB4*01:22 | HG00012948 | HLA15015 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:23 | 100737 | HLA15016 | - | - | - | - | - | - | Dr. Ute Geigenmuller |
| DRB4*01:24 | 070667 | HLA15017 | - | - | - | - | - | - | Dr. Ute Geigenmuller |
| DRB4*01:25 | 074200 | HLA15018 | - | - | - | - | - | - | Dr. Ute Geigenmuller |
| DRB4*01:26 | HG00011751 | HLA15372 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:27 | HG00011752 | HLA15373 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:28 | HG00011753 | HLA15374 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:28 | HG00013044 | HLA15374 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:29 | HG00011754 | HLA15375 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:30 | HG00011756 | HLA15377 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:31 | HG00011757 | HLA15378 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:32 | HG00011758 | HLA15379 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:33 | HG00011759 | HLA15380 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:34 | HG00011761 | HLA15381 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:35 | HG00011762 | HLA15382 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:36 | HG00012475 HG00012476 |
HLA15383 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:36 | HG00011764 HG00011767 HG00011766 HG00011765 |
HLA15383 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:36 | HG00013047 | HLA15383 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:37 | HG00011768 | HLA15429 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:38N | HG00011769 | HLA15430 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:39 | HG00011771 | HLA15431 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:39 | HG00013049 | HLA15431 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:40 | HG00011772 | HLA15432 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:40 | HG00013050 | HLA15432 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:41 | HG00012485 HG00012483 HG00012484 |
HLA15433 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:41 | HG00011773 | HLA15433 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:42 | HG00012202 | HLA15713 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:43 | HG00012203 | HLA15714 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:44 | HG00012480 HG00012482 HG00012479 HG00012478 HG00012481 HG00012477 |
HLA15715 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:44 | HG00013058 HG00013056 HG00013051 HG00013059 HG00013054 HG00013053 HG00013057 HG00013055 HG00013052 |
HLA15715 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:44 | HG00012204 HG00012205 |
HLA15715 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:44 | KW105 | HLA15715 | Jahromi M, Rahma Z, Taha N, Lin PY, Yang KL | Detection of the HLA-DRB4*01:44 allele in a Kuwaiti individual | HLA | 2020 | 96 | 537-539 | Prof. Edward KL Yang |
| DRB4*01:45 | HG00012722 HG00012721 |
HLA16588 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:46 | HG00012724 | HLA16589 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:47 | HG00012725 | HLA16590 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:48 | HG00012727 | HLA16591 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:49 | HG00012862 | HLA16800 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:50 | HG00012950 | HLA16968 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:51 | HG00012951 | HLA16969 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:52 | HG00013178 | HLA17537 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:52 | HG00031358 | HLA17537 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:52 | HG00015931 | HLA17537 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:53 | HG00013179 HG00013180 |
HLA17538 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:54:01N | HG00013181 | HLA17539 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:54:01N | HG00014679 | HLA17539 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:54:02N | HG00027340 | HLA28543 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:55 | HG00013182 | HLA17540 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:56N | HG00013183 | HLA17541 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:57N | HG00013477 | HLA17943 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:57N | HG00014621 | HLA17943 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:58 | HG00013478 | HLA17944 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:58 | HG00031356 | HLA17944 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:58 | HG00014676 | HLA17944 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:59 | HG00013479 | HLA17945 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:60 | HG00013480 | HLA17946 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:60 | HG00026844 | HLA17946 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:60 | HG00031357 | HLA17946 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:60 | HG00015298 HG00013829 |
HLA17946 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:61:01N | HG00013482 | HLA17947 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:61:02N | HG00051229 | HLA38076 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:62 | 81163755 | HLA18068 | - | - | - | - | - | - | Dr. Daniel Fuerst |
| DRB4*01:63 | 81153001 | HLA18085 | - | - | - | - | - | - | Dr. Daniel Fuerst |
| DRB4*01:64 | HG00013939 HG00013940 |
HLA18632 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:65N | HG00013785 | HLA18684 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:65N | HG00018095 | HLA18684 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:65N | HG00015917 | HLA18684 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:66 | HG00014675 | HLA18792 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:67 | HG00014678 | HLA18793 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:68 | HG00014682 | HLA18794 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:69 | HG00013553 | HLA18820 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:70 | HG00017136 | HLA19679 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:70 | HG00019608 | HLA19679 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:70 | HG00022276 | HLA19679 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:71N | HG00017240 | HLA19826 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:72:01 | HG00016340 | HLA20687 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:72:01 | HG00025632 | HLA20687 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:72:01 | HG00022282 | HLA20687 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:72:01 | HG00022284 | HLA20687 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:72:02 | 23111555279HS27822 | HLA39019 | - | - | - | - | - | - | Dr. Cathi Murphey |
| DRB4*01:73 | HG00016341 | HLA20931 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:73 | HG00022269 | HLA20931 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:74 | HG00016609 | HLA20804 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:74 | HG00022263 | HLA20804 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:74 | HG00022287 HG00022300 |
HLA20804 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:74 | HG00022293 | HLA20804 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:75 | HG00015297 HG00018991 HG00018993 HG00018859 HG00019715 HG00015694 |
HLA21033 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:75 | HG00017497 | HLA21033 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:75 | HG00018610 | HLA21033 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:76:01 | HG00017573 | HLA20820 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:76:01 | HG00017505 | HLA20820 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:76:02 | HG00029514 | HLA30928 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:77 | HG00018097 | HLA20749 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:77 | HG00018097 | HLA20749 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:78 | HG00018098 | HLA20625 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:78 | HG00018988 HG00019710 HG00018987 |
HLA20625 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:78 | HG00022294 | HLA20625 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:79 | HG00018100 | HLA20586 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:80N | HG00017501 HG00017500 |
HLA21562 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:80N | HG00017238 HG00017241 |
HLA21562 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:81 | HG00018847 | HLA21227 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:81 | HG00022296 | HLA21227 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:82 | HG00018848 | HLA21568 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:83 | HG00018849 | HLA21459 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:84N | HG00018850 | HLA21216 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:85 | 90P223809 | HLA21905 | - | - | - | - | - | - | Dr. Daniel Fuerst |
| DRB4*01:85 | Ua2018tp2849 | HLA21905 | - | - | - | - | - | - | Dr. Mats Bengtsson |
| DRB4*01:86 | 81192102 | HLA21774 | - | - | - | - | - | - | Dr. Daniel Fuerst |
| DRB4*01:87 | HG00018612 | HLA22622 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:88 | HG00018989 | HLA22631 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:88 | HG00022274 HG00022271 |
HLA22631 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:89 | HG00018990 | HLA22349 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:89 | HG00013828 | HLA22349 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:90 | HG00019610 | HLA22202 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:91 | HG00019711 | HLA22588 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:92 | HG00019712 | HLA22397 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:93 | HG00019713 | HLA22252 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:94 | HG00022298 | HLA23596 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:95 | HG00022301 | HLA23137 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:96 | HG00022289 | HLA23132 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:97 | HG00022290 | HLA23501 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:98 | HG00013554 | HLA23615 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:99 | HG00013825 | HLA23389 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:100 | HG00013826 | HLA23628 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:101 | HG00013827 | HLA23381 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:102 | HG00013830 | HLA23321 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:102 | 114670 | HLA23321 | - | - | - | - | - | - | Dr. Francisco de Paula Sanchez Gordo |
| DRB4*01:103 | HG00022272 | HLA23313 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:104 | HG00022273 | HLA23054 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:105 | HG00022283 | HLA23639 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:106 | HG00016612 HG00015916 |
HLA23452 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:106 | HG00022285 | HLA23452 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:107 | HG00022502 | HLA23438 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:107 | HG00017498 | HLA23438 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:108N | HG00014680 | HLA24284 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:109 | HG00015299 | HLA24228 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:110 | HG00015692 | HLA23912 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:111 | HG00016610 | HLA24130 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:112 | HG00031354 | HLA24266 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:112 | HG00016613 | HLA24266 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:112 | HG00017509 | HLA24266 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:113N | HG00017239 HG00018099 HG00019709 |
HLA24119 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:114 | HG00017511 | HLA23830 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:115N | HG00019607 | HLA24025 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:116 | HG00022487 | HLA23858 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:117 | HG00018617 | HLA24749 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:118 | HG00027345 | HLA24874 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:118 | HG00025060 | HLA24874 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:118 | HG00050360 | HLA24874 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:119 | 6123467 | HLA25855 | - | - | - | - | - | - | Dr. Antonio Balas |
| DRB4*01:119 | HG00029060 | HLA25855 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:119 | HG00025633 | HLA25855 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:119 | 114922 | HLA25855 | - | - | - | - | - | - | Dr. Francisco de Paula Sanchez Gordo |
| DRB4*01:120 | HG00025634 | HLA25994 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:121N | HG00025635 | HLA25805 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:121N | HG00027349 | HLA25805 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:122 | HG00025853 | HLA26299 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:123 | 84028210 | HLA26580 | - | - | - | - | - | - | Dr. Daniel Fuerst |
| DRB4*01:124 | UOU0054 UOU0016 UOU0056 |
HLA26722 | - | - | - | - | - | - | Dr. Eszter Lazar-Molnar |
| DRB4*01:125 | c0546109 | HLA26946 | - | - | - | - | - | - | Dr. Francisco de Paula Sanchez Gordo |
| DRB4*01:126 | HG00027356 | HLA28536 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:127 | HG00029078 | HLA28537 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:127 | HG00029066 HG00029076 HG00029084 |
HLA28537 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:127 | HG00027343 HG00027352 |
HLA28537 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:128N | HG00027344 | HLA28538 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:128N | HG00050362 | HLA28538 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:128N | HG00029511 | HLA28538 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:129 | HG00027357 | HLA28539 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:130 | HG00027358 | HLA28540 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:131 | HG00027347 | HLA28541 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:132 | HG00027348 | HLA28542 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:133 | HG00027355 | HLA28545 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:134 | 20211439 | HLA28680 | - | - | - | - | - | - | Dr. Daniel Fuerst |
| DRB4*01:135 | HG00026845 | HLA28874 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:136 | HG00027350 HG00027353 |
HLA28875 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:136 | HG00026843 | HLA28875 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:137 | HG00026846 | HLA28876 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:138 | HG00027359 HG00029061 HG00029065 HG00029074 |
HLA29155 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:138 | HG00029068 | HLA29155 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:138 | HG00049819 | HLA29155 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:138 | HG00029513 | HLA29155 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:138 | HG00031521 | HLA29155 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:138 | HG00026386 | HLA29155 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:138 | 78820724835 | HLA29155 | - | - | - | - | - | - | Dr. Cathi Murphey |
| DRB4*01:139 | HG00029072 | HLA29839 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:140 | HG00029070 | HLA30269 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:141 | HG00029077 | HLA30386 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:142 | HG00029083 | HLA30343 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:143 | HG00029082 | HLA30378 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:143 | HG00050363 | HLA30378 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:144 | HG00029075 | HLA30430 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:145N | HG00029062 HG00029086 |
HLA30677 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:146 | 21203211 | HLA30601 | - | - | - | - | - | - | Dr. Daniel Fuerst |
| DRB4*01:147 | HG00029508 | HLA31372 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:148 | HG00029515 | HLA31395 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:149N | HG00029714 | HLA31451 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:150 | HG00030143 | HLA31977 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:151 | 2110307078 | HLA32287 | Cargou M, Andreani M, Troiano M, Wojciechowski E, Visentin J | Characterization of the novel HLA-DRB4*01:151 allele by sequencing-based typing. | HLA | 2022 | 99 | 64-66 | Dr. Jonathan Visentin |
| DRB4*01:152 | KUHSC-24-21 | HLA32168 | Ameen R, Geo JA, Thomas J | Identification of a novel HLA-DRB4 allele, HLA-DRB4*01:152 in a Kuwaiti family. | HLA | 2022 | 100 | 546-547 | Dr. Reem Ameen |
| DRB4*01:153 | HG00031359 | HLA32726 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:154 | HG00031360 | HLA32911 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:155 | 2022S94 | HLA34347 | - | - | - | - | - | - | Dr. Alexandre Walencik |
| DRB4*01:156 | 22S140 | HLA34373 | - | - | - | - | - | - | Dr. Alexandre Walencik |
| DRB4*01:157 | HG00031980 | HLA34262 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:158 | HG00027346 | HLA35307 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:159N | HG00015691 | HLA34925 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:160 | HG00050164 HG00049910 |
HLA34958 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:161 | HG00050076 | HLA34898 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:162N | YUMC 006 | HLA35013 | Kim SW, Hong JY, Jung S, Lee J, Park Y | Identification of the novel HLA-DRB4*01:162N allele using next-generation sequencing. | HLA | 2022 | 100 | 659-660 | Dr. Yoonhee Park |
| DRB4*01:163 | HG00050288 | HLA35953 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:164 | 5997073-264317 | HLA36154 | - | - | - | - | - | - | Dr. Neifi Hassan Saloum Deghaide |
| DRB4*01:165N | HILAPL-001 | HLA36512 | Hu Q, Burns C, McKinley C, Christian D, Berka N | Identification of a novel allele with a frameshift mutation, HLA-DRB4*01:165N, using next-generation sequencing. | HLA | 2023 | 101 | 699-700 | Dr. Qian Hu |
| DRB4*01:166 | HG00050547 | HLA36453 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:167 | 5996042-263676 | HLA36603 | - | - | - | - | - | - | Dr. Neifi Hassan Saloum Deghaide |
| DRB4*01:168 | 22S181 | HLA36964 | - | - | - | - | - | - | Dr. Alexandre Walencik |
| DRB4*01:169 | TPH22-49-F2 | HLA37076 | - | - | - | - | - | - | Dr. Antonio Nieto |
| DRB4*01:170 | HG00050515 | HLA37332 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:171 | HG00050948 | HLA37293 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:172 | 81279727 | HLA37189 | - | - | - | - | - | - | Dr. Daniel Fuerst |
| DRB4*01:173 | 23002272 | HLA38034 | - | - | - | - | - | - | Dr. Robert Bray |
| DRB4*01:174 | HG00051230 | HLA38073 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:175 | 521/23 | HLA38471 | - | - | - | - | - | - | Mr. Rafael Oliveira |
| DRB4*01:176 | 139459821 | HLA38466 | - | - | - | - | - | - | Dr. Pascal Pedini |
| DRB4*01:177 | HG00051309 | HLA38352 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:178 | HG00051424 | HLA38705 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:179 | SZ-213 | HLA39346 | Wu WD, Zhong XX, Quan ZR, Luo SH, Yuan R | A novel HLA-DRB4 allele, HLA-DRB4*01:179. | HLA | 2024 | 104 | - | Dr. Hong-Yan Zou |
| DRB4*01:180 | 24202090 | HLA39729 | - | - | - | - | - | - | Dr. Daniel Fuerst |
| DRB4*01:181 | HG00051934 | HLA40065 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:182 | 2400402 | HLA40299 | Blandin L, Cargou M, Wojciechowski E, Lemal R, Rouzaire P | Characterisation of the novel HLA-DRB4*01:182 allele by sequencing-based typing. | HLA | 2024 | 104 | - | Dr. Paul Rouzaire |
| DRB4*01:183 | JH0125 | HLA40783 | - | - | - | - | - | - | Dr. Maria Bettinotti |
| DRB4*01:184 | HG00052159 | HLA40897 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:185 | HG00052192 | HLA40940 | - | - | - | - | - | - | Dr. Nezih Cereb |
| DRB4*01:186 | FSIMM_IMM-23-81685 | HLA41096 | - | - | - | - | - | - | Dr. Dianne De Santis |
| DRB4*01:187 | RP-285737 | HLA41270 | - | - | - | - | - | - | Dr. Neifi Hassan Saloum Deghaide |
| DRB4*01:188 | 20QR006369 | HLA41503 | - | - | - | - | - | - | Dr. Janette Kwok |
| DRB4*02:01N | GN016 | HLA00913 | Robbins F, Hurley CK, Tang T, Yao H, Lin YS, Wade J, Goeken N, Hartzman RJ | Diversity associated with the second expressed HLA-DRB locus in the human population | Immunogenetics | 1997 | 46 | 104-110 | Dr. Fu-Meei Robbins |
| DRB4*02:01N | GN016 | HLA00913 | - | - | - | - | - | - | Prof. Carolyn K Hurley |
| DRB4*03:01N | GN017 | HLA00914 | Robbins F, Hurley CK, Tang T, Yao H, Lin YS, Wade J, Goeken N, Hartzman RJ | Diversity associated with the second expressed HLA-DRB locus in the human population | Immunogenetics | 1997 | 46 | 104-110 | Dr. Fu-Meei Robbins |
| DRB4*03:01N | GN017 GN020 GN076 GN092 |
HLA00914 | Steiner NK, Hou L, Hurley CK | Characterizing allele with large deletions using region specific extraction | Human Immunology | 2018 | 79 | 491-493 | Prof. Carolyn K Hurley |
