DRB4 References

Assigned March 2024

Allele Name Cell ID HLA ID Authors Title Journal Year Vol. Pages Contact
DRB4*01:01:01:01 FS
LKT3		 HLA00905 Gregersen PK, Shen M, Song QL, Merryman P, Degar S, Seki T, Maccari J, Goldberg D, Murphy H, Schwenzer J, Wang CY, Winchester RJ, Nepom GT, Silver J Molecular diversity of HLA-DR4 haplotypes Proceedings of the National Academy of Sciences USA 1986 83 2642-2646 Dr. Peter Gregersen
DRB4*01:01:01:01 BURKHARDT HLA00905 Gregersen PK, Moriuchi T, Karr RW, Obata F, Moriuchi J, Maccari J, Goldberg D, Winchester RJ, Silver J Polymorphism of HLA-DR beta chains in DR4, -7, and -9 haplotypes: implications for the mechanisms of allelic variation Proceedings of the National Academy of Sciences USA 1986 83 9149-9153 Dr. Peter Gregersen
DRB4*01:01:01:01 PRIESS HLA00905 Spies T, Sorrentino R, Boss JM, Okada K, Strominger JL Structural organization of the DR subregion of the human major histocompatibility complex Proceedings of the National Academy of Sciences USA 1985 82 5165-5169 Prof. Jack L Strominger
DRB4*01:01:01:01 DM24
DM29		 HLA00905 Todd JA, Bell JI, McDevitt HO HLA-DQ beta gene contributes to susceptibility and resistance to insulin-dependent diabetes mellitus Nature 1987 329 599-604 Dr. John A. Todd
DRB4*01:01:01:01 MMCC HLA00905 Horn GT, Bugawan TL, Long CM, Manos MM, Erlich HA Sequence analysis of HLA class II genes from insulin-dependent diabetic individuals Human Immunology 1988 21 249-263 Dr. Henry Erlich
DRB4*01:01:01:01 MOU HLA00905 Young JA, Wilkinson D, Bodmer WF, Trowsdale J Sequence and evolution of HLA-DR7- and -DRw53-associated beta-chain genes Proceedings of the National Academy of Sciences USA 1987 84 4929-4933 Prof. John Trowsdale
DRB4*01:01:01:01 MOU HLA00905 Horton R, Gibson R, Coggill P, Miretti M, Allcok RJ, Almeida J, Forbes S, Gilbert JGR, Halls K, Harrow JL, Hart E, Howe K, Jackson DK, Palmer S, Roberts AN, Sims S, Stewart CA, Traherne JA, Trevanion S, Wilming L, Rogers J, de Jong PJ, Elliott JF, Sawcer S, Todd JA, Trowsdale J, Beck S Variation analysis and gene annotation of eight MHC haplotypes: The MHC Haplotype Project. Immunogenetics 2008 60 1-18 Prof. Stephan Beck
DRB4*01:01:01:01 MOLT-4 HLA00905 Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. HLA 2022 99 328-356 Prof. Steven GE Marsh
DRB4*01:01:01:01 FH05 HLA00905 Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. HLA 2022 99 328-356 Prof. Steven GE Marsh
DRB4*01:01:01:03 ARBO HLA23372 Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. HLA 2022 99 328-356 Prof. Steven GE Marsh
DRB4*01:01:01:04 SHJO HLA31732 Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. HLA 2022 99 328-356 Prof. Steven GE Marsh
DRB4*01:01:02 HG00011755 HLA15376 - - - - - - Dr. Nezih Cereb
DRB4*01:01:02 HG00012473 HLA15376 - - - - - - Dr. Nezih Cereb
DRB4*01:01:03 HG00011749
HG00011750		 HLA22240 - - - - - - Dr. Nezih Cereb
DRB4*01:01:03 HG00018101 HLA22240 - - - - - - Dr. Nezih Cereb
DRB4*01:01:03 HG00022266 HLA22240 - - - - - - Dr. Nezih Cereb
DRB4*01:01:04 PN39817732AN HLA22373 - - - - - - Prof. Steven GE Marsh
DRB4*01:01:04 HG00011286 HLA22373 - - - - - - Dr. Nezih Cereb
DRB4*01:01:04 HG00018613 HLA22373 - - - - - - Dr. Nezih Cereb
DRB4*01:01:04 HG00022292 HLA22373 - - - - - - Dr. Nezih Cereb
DRB4*01:01:04 HG00025623
HG00025624		 HLA22373 - - - - - - Dr. Nezih Cereb
DRB4*01:01:04 HG00025376 HLA22373 - - - - - - Dr. Nezih Cereb
DRB4*01:01:04 HG00014619
HG00014681
HG00014683
HG00015913
HG00015914
HG00016608
HG00018103
HG00018618
HG00022275
HG00022291		 HLA22373 - - - - - - Dr. Nezih Cereb
DRB4*01:01:04 c2128591 HLA22373 - - - - - - Dr. Francisco de Paula Sanchez Gordo
DRB4*01:01:05 HG00014618 HLA22344 - - - - - - Dr. Nezih Cereb
DRB4*01:01:05 HG00018611 HLA22344 - - - - - - Dr. Nezih Cereb
DRB4*01:01:06 81196967 HLA22452 - - - - - - Dr. Daniel Fuerst
DRB4*01:01:07 HG00012949 HLA23515 - - - - - - Dr. Nezih Cereb
DRB4*01:01:07 HG00022267 HLA23515 - - - - - - Dr. Nezih Cereb
DRB4*01:01:07 HG00019609 HLA23515 - - - - - - Dr. Nezih Cereb
DRB4*01:01:08 HG00022286 HLA23595 - - - - - - Dr. Nezih Cereb
DRB4*01:01:09 HG00022561 HLA25824 - - - - - - Dr. Nezih Cereb
DRB4*01:01:09 HG00027341 HLA25824 - - - - - - Dr. Nezih Cereb
DRB4*01:01:10 HG00029058 HLA30109 - - - - - - Dr. Nezih Cereb
DRB4*01:01:10 HG00031774 HLA30109 - - - - - - Dr. Nezih Cereb
DRB4*01:01:11 23032847726HS2680 HLA37518 - - - - - - Dr. Cathi Murphey
DRB4*01:01:11 HS2680 HLA37518 - - - - - - Dr. Cathi Murphey
DRB4*01:01:12 2210491407 HLA37807 - - - - - - Dr. Jonathan Visentin
DRB4*01:01:13 P14847-1 HLA39861 - - - - - - Prof. Faming Zhu
DRB4*01:02 C.M.L. HLA00907 Naruse TK, Ando R, Nose Y, Kagiya M, Ando H, Kawata H, Nabeya N, Isshiki G, Inoko H HLA-DRB4 genotyping by PCR-RFLP: diversity in the associations between HLA-DRB4 and DRB1 alleles Tissue Antigens 1997 49 152-159 Prof. Hidetoshi Inoko
DRB4*01:02 C.M.L. HLA00907 Buyse I, Emonds MP, Bouillon R, Marynen P, Cassiman JJ Novel class II HLA-DRB4 and DPB1 alleles found in the Belgian population Immunogenetics 1993 38 380- Prof. Jean-Jacques Cassiman
DRB4*01:02 C.M.L. HLA00907 Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. HLA 2022 99 328-356 Prof. Steven GE Marsh
DRB4*01:03:01:01 MJ4 HLA00908 Curtsinger JM, Hilden JM, Cairns JS, Bach FH Evolutionary and genetic implications of sequence variation in two nonallelic HLA-DR beta-chain cDNA sequences Proceedings of the National Academy of Sciences USA 1987 84 209-213 Prof. Fritz H Bach
DRB4*01:03:01:01 HSF7 HLA00908 - - - - - - Prof. Stephan Beck
DRB4*01:03:01:01 G081 HLA00908 De Pablo R, Solis R, Balas A, Vilches C Specific amplification of the HLA-DRB4 gene from c-DNA. Complete coding sequence of the HLA alleles DRB4*0103101 and DRB4*01033 Tissue Antigens 2002 59 44-46 Dr. Rosario de Pablo
DRB4*01:03:01:01 MLF HLA00908 Horton R, Gibson R, Coggill P, Miretti M, Allcok RJ, Almeida J, Forbes S, Gilbert JGR, Halls K, Harrow JL, Hart E, Howe K, Jackson DK, Palmer S, Roberts AN, Sims S, Stewart CA, Traherne JA, Trevanion S, Wilming L, Rogers J, de Jong PJ, Elliott JF, Sawcer S, Todd JA, Trowsdale J, Beck S Variation analysis and gene annotation of eight MHC haplotypes: The MHC Haplotype Project. Immunogenetics 2008 60 1-18 Prof. Stephan Beck
DRB4*01:03:01:02N DBB HLA00909 Sutton VR, Knowles RW An aberrant DRB4 null gene transcript is found that could encode a novel HLA-DR beta chain Immunogenetics 1990 31 112-117 Dr. Robert W Knowles
DRB4*01:03:01:02N DBB HLA00909 Naruse TK, Ando R, Nose Y, Kagiya M, Ando H, Kawata H, Nabeya N, Isshiki G, Inoko H HLA-DRB4 genotyping by PCR-RFLP: diversity in the associations between HLA-DRB4 and DRB1 alleles Tissue Antigens 1997 49 152-159 Prof. Hidetoshi Inoko
DRB4*01:03:01:02N DBB HLA00909 Sutton VR, Knowles RW An aberrant DRB4 null gene transcript is found that could encode a novel HLA-DR beta chain Immunogenetics 1990 31 112-117 Dr. Robert W Knowles
DRB4*01:03:01:02N MYE 2004 HLA00909 - - - - - - Mr. Konstantinos Barsakis
DRB4*01:03:01:02N BY01567
BY01568		 HLA00909 - - - - - - Prof. Carolyn K Hurley
DRB4*01:03:01:02N DBB HLA00909 - - - - - - Prof. Carolyn K Hurley
DRB4*01:03:01:02N DBB HLA00909 Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. HLA 2022 99 328-356 Prof. Steven GE Marsh
DRB4*01:03:01:02N MYE 2004 HLA00909 Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. HLA 2022 99 328-356 Prof. Steven GE Marsh
DRB4*01:03:01:02N WIN HLA00909 Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. HLA 2022 99 328-356 Prof. Steven GE Marsh
DRB4*01:03:01:03 SSTO HLA06603 Horton R, Gibson R, Coggill P, Miretti M, Allcok RJ, Almeida J, Forbes S, Gilbert JGR, Halls K, Harrow JL, Hart E, Howe K, Jackson DK, Palmer S, Roberts AN, Sims S, Stewart CA, Traherne JA, Trevanion S, Wilming L, Rogers J, de Jong PJ, Elliott JF, Sawcer S, Todd JA, Trowsdale J, Beck S Variation analysis and gene annotation of eight MHC haplotypes: The MHC Haplotype Project. Immunogenetics 2008 60 1-18 Prof. Stephan Beck
DRB4*01:03:01:03 SSTO HLA06603 Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. HLA 2022 99 328-356 Prof. Steven GE Marsh
DRB4*01:03:01:03 FH12 HLA06603 Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. HLA 2022 99 328-356 Prof. Steven GE Marsh
DRB4*01:03:01:04 LBF
LBUF		 HLA20773 Bell JI, Denney D Jr, Foster L, Belt T, Todd JA, McDevitt HO Allelic variation in the DR subregion of the human major histocompatibility complex Proceedings of the National Academy of Sciences USA 1987 84 6234-6248 Prof. John Bell
DRB4*01:03:01:04 SHJO HLA20773 Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. HLA 2022 99 328-356 Prof. Steven GE Marsh
DRB4*01:03:01:04 EAV, AC HLA20773 Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. HLA 2022 99 328-356 Prof. Steven GE Marsh
DRB4*01:03:01:04 BH HLA20773 Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. HLA 2022 99 328-356 Prof. Steven GE Marsh
DRB4*01:03:01:04 LBF
LBUF		 HLA20773 - - - - - - Prof. Steven GE Marsh
DRB4*01:03:01:05 LKT3 HLA20628 Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. HLA 2022 99 328-356 Prof. Steven GE Marsh
DRB4*01:03:01:05 KY HLA20628 Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. HLA 2022 99 328-356 Prof. Steven GE Marsh
DRB4*01:03:01:06 SYSU005 HLA21590 - - - - - - Dr. Riga Wu
DRB4*01:03:01:07 CHI 007 HLA21306 Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. HLA 2022 99 328-356 Prof. Steven GE Marsh
DRB4*01:03:01:08 T7527 HLA21599 Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. HLA 2022 99 328-356 Prof. Steven GE Marsh
DRB4*01:03:01:09 DEU HLA22658 Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. HLA 2022 99 328-356 Prof. Steven GE Marsh
DRB4*01:03:01:10 YAR HLA22663 Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. HLA 2022 99 328-356 Prof. Steven GE Marsh
DRB4*01:03:01:11 BOLETH HLA22666 Andersson G, Larhammar D, Widmark E, Servenius B, Peterson PA, Rask L Class II genes of the human major histocompatibility complex. Organization and evolutionary relationship of the DR beta genes Journal of Biological Chemistry 1987 262 8748-8758 Dr. Goran Andersson
DRB4*01:03:01:11 BM14 HLA22666 Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. HLA 2022 99 328-356 Prof. Steven GE Marsh
DRB4*01:03:01:11 JHAF HLA22666 Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. HLA 2022 99 328-356 Prof. Steven GE Marsh
DRB4*01:03:01:11 BOLETH HLA22666 - - - - - - Prof. Steven GE Marsh
DRB4*01:03:01:12 BUR, E HLA23081 Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. HLA 2022 99 328-356 Prof. Steven GE Marsh
DRB4*01:03:01:13N HB19 HLA26755 - - - - - - Dr. Riga Wu
DRB4*01:03:01:14 MANIKA HLA29115 Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. HLA 2022 99 328-356 Prof. Steven GE Marsh
DRB4*01:03:01:15 FB11 HLA30796 Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. HLA 2022 99 328-356 Prof. Steven GE Marsh
DRB4*01:03:01:16 BH HLA31733 Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. HLA 2022 99 328-356 Prof. Steven GE Marsh
DRB4*01:03:01:17 EC HLA31515 Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. HLA 2022 99 328-356 Prof. Steven GE Marsh
DRB4*01:03:01:18 AN302594 HLA38110 - - - - - - Prof. Steven GE Marsh
DRB4*01:03:02:01 DKB HLA00910 Bell JI, Denney D Jr, Foster L, Belt T, Todd JA, McDevitt HO Allelic variation in the DR subregion of the human major histocompatibility complex Proceedings of the National Academy of Sciences USA 1987 84 6234-6248 Prof. John Bell
DRB4*01:03:02:01 W778R HLA00910 - - - - - - Prof. Lee Ann Baxter-Lowe
DRB4*01:03:02:01 FKD4010302E1 HLA00910 - - - - - - Mr. Konstantinos Barsakis
DRB4*01:03:02:01 DKB HLA00910 Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. HLA 2022 99 328-356 Prof. Steven GE Marsh
DRB4*01:03:02:01 LKT14 HLA00910 Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. HLA 2022 99 328-356 Prof. Steven GE Marsh
DRB4*01:03:02:02 GRC-212 HLA31794 Turner TR, Hayward DR, Gymer AW, Barker DJ, Leen G, Cambridge CA, Macpherson HL, Georgiou X, Cooper MA, Robinson J, Mayor NP, Marsh SGE Widespread non-coding polymorphism in HLA class II genes of International HLA and Immunogenetics Workshop cell lines. HLA 2022 99 328-356 Prof. Steven GE Marsh
DRB4*01:03:03 MG-CV HLA01225 Voorter CE, Hepkema BG, Lems SP, van den Berg-Loonen EM Identification of three new DRB1 alleles, DRB1*0107, *0425 and *13012 and confirmation of DRB4*01033 Tissue Antigens 2003 61 398-402 Prof. Ella M. Van den Berg-Loonen
DRB4*01:03:03 FOA2362 HLA01225 Palou E, Nogues N, Gil J, Ribera A Identification of a new HLA-DRB4 allele (DRB4*01033) by PCR-SSP and direct sequencing Tissue Antigens 2000 56 279-281 Dr. Eduard Palou
DRB4*01:03:03 MUDGH HLA01225 - - - - - - Prof. Gottfried Fischer
DRB4*01:03:03 G081 HLA01225 De Pablo R, Solis R, Balas A, Vilches C Specific amplification of the HLA-DRB4 gene from c-DNA. Complete coding sequence of the HLA alleles DRB4*0103101 and DRB4*01033 Tissue Antigens 2002 59 44-46 Dr. Rosario de Pablo
DRB4*01:03:04 14242 HLA01594 Fasano ME, Dametto E, Day S, Dunn P, Tacconella M, Curtoni ES A new allele, HLA-DRB4*010304 Tissue Antigens 2003 62 76-78 Dr. Maria Edvige Fasano
DRB4*01:03:05 113757 HLA17334 - - - - - - Dr. Ute Geigenmuller
DRB4*01:03:05 HG00012723 HLA17334 - - - - - - Dr. Nezih Cereb
DRB4*01:03:05 HG00031350 HLA17334 - - - - - - Dr. Nezih Cereb
DRB4*01:03:06 81172337 HLA18069 - - - - - - Dr. Daniel Fuerst
DRB4*01:03:07 HG00011760 HLA19800 - - - - - - Dr. Nezih Cereb
DRB4*01:03:07 HG00017244 HLA19800 - - - - - - Dr. Nezih Cereb
DRB4*01:03:07 HG00017502
HG00017504		 HLA19800 - - - - - - Dr. Nezih Cereb
DRB4*01:03:07 HG00022262
HG00022281		 HLA19800 - - - - - - Dr. Nezih Cereb
DRB4*01:03:07 HG00022279 HLA19800 - - - - - - Dr. Nezih Cereb
DRB4*01:03:07 HG00013831 HLA19800 - - - - - - Dr. Nezih Cereb
DRB4*01:03:07 HG00017503
HG00017508		 HLA19800 - - - - - - Dr. Nezih Cereb
DRB4*01:03:07 HG00018096 HLA19800 - - - - - - Dr. Nezih Cereb
DRB4*01:03:07 HG00025630 HLA19800 - - - - - - Dr. Nezih Cereb
DRB4*01:03:07 HG00031349 HLA19800 - - - - - - Dr. Nezih Cereb
DRB4*01:03:08 HG00017506 HLA20967 - - - - - - Dr. Nezih Cereb
DRB4*01:03:08 HG00031355 HLA20967 - - - - - - Dr. Nezih Cereb
DRB4*01:03:09 HG00013481 HLA20759 - - - - - - Dr. Nezih Cereb
DRB4*01:03:09 HG00017507 HLA20759 - - - - - - Dr. Nezih Cereb
DRB4*01:03:09 HG00019054 HLA20759 - - - - - - Dr. Nezih Cereb
DRB4*01:03:09 HG00022295 HLA20759 - - - - - - Dr. Nezih Cereb
DRB4*01:03:09 HG00022299 HLA20759 - - - - - - Dr. Nezih Cereb
DRB4*01:03:09 HG00015915
HG00016901
HG00017235
HG00018851		 HLA20759 - - - - - - Dr. Nezih Cereb
DRB4*01:03:09 HG00022261 HLA20759 - - - - - - Dr. Nezih Cereb
DRB4*01:03:09 HG00022280 HLA20759 - - - - - - Dr. Nezih Cereb
DRB4*01:03:09 HG00025622 HLA20759 - - - - - - Dr. Nezih Cereb
DRB4*01:03:10 HG00018102 HLA21092 - - - - - - Dr. Nezih Cereb
DRB4*01:03:11 HG00012201 HLA21543 - - - - - - Dr. Nezih Cereb
DRB4*01:03:11 HG00018852 HLA21543 - - - - - - Dr. Nezih Cereb
DRB4*01:03:11 HG00022268 HLA21543 - - - - - - Dr. Nezih Cereb
DRB4*01:03:12 HG00013483 HLA23180 - - - - - - Dr. Nezih Cereb
DRB4*01:03:12 HG00022297 HLA23180 - - - - - - Dr. Nezih Cereb
DRB4*01:03:13 HG00017510 HLA23123 - - - - - - Dr. Nezih Cereb
DRB4*01:03:13 HG00018614
HG00018616
HG00018853
HG00018854
HG00018855
HG00018857		 HLA23123 - - - - - - Dr. Nezih Cereb
DRB4*01:03:13 HG00018856 HLA23123 - - - - - - Dr. Nezih Cereb
DRB4*01:03:13 HG00025375
HG00025627
HG00025629		 HLA23123 - - - - - - Dr. Nezih Cereb
DRB4*01:03:13 HG00025625 HLA23123 - - - - - - Dr. Nezih Cereb
DRB4*01:03:13 HG00025628 HLA23123 - - - - - - Dr. Nezih Cereb
DRB4*01:03:13 HG00013824
HG00015296
HG00016902		 HLA23123 - - - - - - Dr. Nezih Cereb
DRB4*01:03:13 HG00018615 HLA23123 - - - - - - Dr. Nezih Cereb
DRB4*01:03:14 HG00013832 HLA23087 - - - - - - Dr. Nezih Cereb
DRB4*01:03:14 HG00017512 HLA23087 - - - - - - Dr. Nezih Cereb
DRB4*01:03:15 HG00022277 HLA24064 - - - - - - Dr. Nezih Cereb
DRB4*01:03:16 HG00011770 HLA24498 - - - - - - Dr. Nezih Cereb
DRB4*01:03:16 HG00024988 HLA24498 - - - - - - Dr. Nezih Cereb
DRB4*01:03:16 HG00027354 HLA24498 - - - - - - Dr. Nezih Cereb
DRB4*01:03:17 HG00025061 HLA25538 - - - - - - Dr. Nezih Cereb
DRB4*01:03:17 HG00025852 HLA25538 - - - - - - Dr. Nezih Cereb
DRB4*01:03:17 HG00029085 HLA25538 - - - - - - Dr. Nezih Cereb
DRB4*01:03:18 HG00025631 HLA25916 - - - - - - Dr. Nezih Cereb
DRB4*01:03:19 HG00026852 HLA28544 - - - - - - Dr. Nezih Cereb
DRB4*01:03:20 GSQ-58 HLA29793 - - - - - - Prof. Su-Qing Gao
DRB4*01:03:21 HG00029063 HLA30481 - - - - - - Dr. Nezih Cereb
DRB4*01:03:22 HG00030015 HLA31560 - - - - - - Dr. Nezih Cereb
DRB4*01:03:23 HG00017236 HLA31941 - - - - - - Dr. Nezih Cereb
DRB4*01:03:23 TUJM336 HLA31941 - - - - - - Prof. Takashi Shiina
DRB4*01:03:24 HG00030145 HLA31976 - - - - - - Dr. Nezih Cereb
DRB4*01:03:25 HG00030144 HLA31991 - - - - - - Dr. Nezih Cereb
DRB4*01:03:26 HG00011289 HLA34369 - - - - - - Dr. Nezih Cereb
DRB4*01:03:26 HG00022288 HLA34369 - - - - - - Dr. Nezih Cereb
DRB4*01:03:26 HG00031981 HLA34369 - - - - - - Dr. Nezih Cereb
DRB4*01:03:26 HG00013938 HLA34369 - - - - - - Dr. Nezih Cereb
DRB4*01:03:27 HG00015695 HLA34900 - - - - - - Dr. Nezih Cereb
DRB4*01:03:27 HG00031982 HLA34900 - - - - - - Dr. Nezih Cereb
DRB4*01:03:28 HG00050633 HLA36615 - - - - - - Dr. Nezih Cereb
DRB4*01:03:29 HG00050476 HLA36797 - - - - - - Dr. Nezih Cereb
DRB4*01:03:30 922900123 HLA36817 - - - - - - Dr. Dolores Planelles
DRB4*01:03:31 HG00050768 HLA36889 - - - - - - Dr. Nezih Cereb
DRB4*01:03:32 HG00050859
HG00050860		 HLA37099 - - - - - - Dr. Nezih Cereb
DRB4*01:03:33 23032147481HS26773 HLA37511 - - - - - - Dr. Cathi Murphey
DRB4*01:03:33 HS26773HK HLA37511 - - - - - - Dr. Cathi Murphey
DRB4*01:03:34 HG00051164 HLA37796 - - - - - - Dr. Nezih Cereb
DRB4*01:03:35 20029650 HLA38781 - - - - - - Dr. Silvia Giuliodori
DRB4*01:04 69-218
76-394		 HLA00911 De Canck I, Demanet C, Mersch G, Jannes G, Rombout A, Rossau R, Van Waeyenberge C Characterization of a new DRB4 allele (DRB4*0104) Tissue Antigens 1996 48 213-216 Dr. Guy Mersch
DRB4*01:04 TBC-4975 HLA00911 - - - - - - Dr. Masahiro Satake
DRB4*01:05 17345 HLA00912 Voorter CE, Emonds MP, van den Berg-Loonen EM Identification of a new DRB4 allele (DRB4*0105) by sequence-based typing Tissue Antigens 1997 49 662-664 Prof. Ella M. Van den Berg-Loonen
DRB4*01:06 MKOST HLA01559 - - - - - - Dr. Heather Dunckley
DRB4*01:07:01 X0002601
X0002630
X0002652		 HLA01891 - - - - - - Dr. Lia Mele
DRB4*01:07:02 122192 HLA15019 - - - - - - Dr. Ute Geigenmuller
DRB4*01:07:02 HG00011763 HLA15019 - - - - - - Dr. Nezih Cereb
DRB4*01:07:02 HG00013045
HG00013046		 HLA15019 - - - - - - Dr. Nezih Cereb
DRB4*01:08 Tokushima HLA04572 Lee E, Ozaki S, Ishii H, Takimoto T, Matsumoto T A novel allele, HLA-DRB1*01:08, identified by sequence-based typing Tissue Antigens 2011 78 158-159 Ms. Etsukp Lee
DRB4*01:09 LUMC-DRB4-100 HLA13511 - - - - - - Miss. Jacqy DH Anholts
DRB4*01:10 LUMC-DRB4-114 HLA14114 - - - - - - Miss. Jacqy DH Anholts
DRB4*01:11 109551 HLA14659 - - - - - - Dr. Ute Geigenmuller
DRB4*01:11 HG00011278 HLA14659 - - - - - - Dr. Nezih Cereb
DRB4*01:11 HG00011972 HLA14659 - - - - - - Dr. Nezih Cereb
DRB4*01:12 HG00011280 HLA14660 - - - - - - Dr. Nezih Cereb
DRB4*01:13 HG00011282 HLA14661 - - - - - - Dr. Nezih Cereb
DRB4*01:13 HG00011973 HLA14661 - - - - - - Dr. Nezih Cereb
DRB4*01:13 HG00012474 HLA14661 - - - - - - Dr. Nezih Cereb
DRB4*01:14N 93053764 HLA14662 - - - - - - Dr. Daniel Fuerst
DRB4*01:14N HG00011283
HG00011284
HG00011285		 HLA14662 - - - - - - Dr. Nezih Cereb
DRB4*01:14N HG00011974
HG00011975
HG00011976
HG00011977
HG00011978
HG00011979
HG00011980
HG00011981
HG00011982
HG00011983		 HLA14662 - - - - - - Dr. Nezih Cereb
DRB4*01:14N 19205151 HLA14662 Fuerst D, Tsamadou C, Gowdavally S, Schrezenmeier H, Mytilineos J HLA-DRB4*01:14 is a null allele, and renamed HLA-DRB4*01:14N HLA 2020 95 73-75 Dr. Daniel Fuerst
DRB4*01:15 HG00011287 HLA14663 - - - - - - Dr. Nezih Cereb
DRB4*01:16N 076099 HLA14664 - - - - - - Dr. Ute Geigenmuller
DRB4*01:16N HG00011288 HLA14664 - - - - - - Dr. Nezih Cereb
DRB4*01:16N HG00013048 HLA14664 - - - - - - Dr. Nezih Cereb
DRB4*01:16N HG00014705
HG00014706		 HLA14664 - - - - - - Dr. Nezih Cereb
DRB4*01:17 HG00011290
HG00011291		 HLA14665 - - - - - - Dr. Nezih Cereb
DRB4*01:17 HG00011984
HG00011985
HG00011986
HG00011987
HG00011988
HG00011989
HG00011990
HG00011991
HG00011992
HG00011993
HG00011994
HG00011995
HG00011996		 HLA14665 - - - - - - Dr. Nezih Cereb
DRB4*01:17 HG00012486
HG00012487		 HLA14665 - - - - - - Dr. Nezih Cereb
DRB4*01:17 HG00013060 HLA14665 - - - - - - Dr. Nezih Cereb
DRB4*01:17 HG00015814
HG00015815		 HLA14665 - - - - - - Dr. Nezih Cereb
DRB4*01:18 HG00011500 HLA15089 - - - - - - Dr. Nezih Cereb
DRB4*01:19 HG00011501 HLA15090 - - - - - - Dr. Nezih Cereb
DRB4*01:20 HG00011502 HLA15091 - - - - - - Dr. Nezih Cereb
DRB4*01:21 HG00011503 HLA15092 - - - - - - Dr. Nezih Cereb
DRB4*01:22 046670 HLA15015 - - - - - - Dr. Ute Geigenmuller
DRB4*01:22 HG00012948 HLA15015 - - - - - - Dr. Nezih Cereb
DRB4*01:23 100737 HLA15016 - - - - - - Dr. Ute Geigenmuller
DRB4*01:24 070667 HLA15017 - - - - - - Dr. Ute Geigenmuller
DRB4*01:25 074200 HLA15018 - - - - - - Dr. Ute Geigenmuller
DRB4*01:26 HG00011751 HLA15372 - - - - - - Dr. Nezih Cereb
DRB4*01:27 HG00011752 HLA15373 - - - - - - Dr. Nezih Cereb
DRB4*01:28 HG00011753 HLA15374 - - - - - - Dr. Nezih Cereb
DRB4*01:28 HG00013044 HLA15374 - - - - - - Dr. Nezih Cereb
DRB4*01:29 HG00011754 HLA15375 - - - - - - Dr. Nezih Cereb
DRB4*01:30 HG00011756 HLA15377 - - - - - - Dr. Nezih Cereb
DRB4*01:31 HG00011757 HLA15378 - - - - - - Dr. Nezih Cereb
DRB4*01:32 HG00011758 HLA15379 - - - - - - Dr. Nezih Cereb
DRB4*01:33 HG00011759 HLA15380 - - - - - - Dr. Nezih Cereb
DRB4*01:34 HG00011761 HLA15381 - - - - - - Dr. Nezih Cereb
DRB4*01:35 HG00011762 HLA15382 - - - - - - Dr. Nezih Cereb
DRB4*01:36 HG00011764
HG00011765
HG00011766
HG00011767		 HLA15383 - - - - - - Dr. Nezih Cereb
DRB4*01:36 HG00012475
HG00012476		 HLA15383 - - - - - - Dr. Nezih Cereb
DRB4*01:36 HG00013047 HLA15383 - - - - - - Dr. Nezih Cereb
DRB4*01:37 HG00011768 HLA15429 - - - - - - Dr. Nezih Cereb
DRB4*01:38N HG00011769 HLA15430 - - - - - - Dr. Nezih Cereb
DRB4*01:39 HG00011771 HLA15431 - - - - - - Dr. Nezih Cereb
DRB4*01:39 HG00013049 HLA15431 - - - - - - Dr. Nezih Cereb
DRB4*01:40 HG00011772 HLA15432 - - - - - - Dr. Nezih Cereb
DRB4*01:40 HG00013050 HLA15432 - - - - - - Dr. Nezih Cereb
DRB4*01:41 HG00011773 HLA15433 - - - - - - Dr. Nezih Cereb
DRB4*01:41 HG00012483
HG00012484
HG00012485		 HLA15433 - - - - - - Dr. Nezih Cereb
DRB4*01:42 HG00012202 HLA15713 - - - - - - Dr. Nezih Cereb
DRB4*01:43 HG00012203 HLA15714 - - - - - - Dr. Nezih Cereb
DRB4*01:44 HG00012204
HG00012205		 HLA15715 - - - - - - Dr. Nezih Cereb
DRB4*01:44 HG00012477
HG00012478
HG00012479
HG00012480
HG00012481
HG00012482		 HLA15715 - - - - - - Dr. Nezih Cereb
DRB4*01:44 HG00013051
HG00013052
HG00013053
HG00013054
HG00013055
HG00013056
HG00013057
HG00013058
HG00013059		 HLA15715 - - - - - - Dr. Nezih Cereb
DRB4*01:44 KW105 HLA15715 Jahromi M, Rahma Z, Taha N, Lin PY, Yang KL Detection of the HLA-DRB4*01:44 allele in a Kuwaiti individual HLA 2020 96 537-539 Prof. Edward KL Yang
DRB4*01:45 HG00012721
HG00012722		 HLA16588 - - - - - - Dr. Nezih Cereb
DRB4*01:46 HG00012724 HLA16589 - - - - - - Dr. Nezih Cereb
DRB4*01:47 HG00012725 HLA16590 - - - - - - Dr. Nezih Cereb
DRB4*01:48 HG00012727 HLA16591 - - - - - - Dr. Nezih Cereb
DRB4*01:49 HG00012862 HLA16800 - - - - - - Dr. Nezih Cereb
DRB4*01:50 HG00012950 HLA16968 - - - - - - Dr. Nezih Cereb
DRB4*01:51 HG00012951 HLA16969 - - - - - - Dr. Nezih Cereb
DRB4*01:52 HG00013178 HLA17537 - - - - - - Dr. Nezih Cereb
DRB4*01:52 HG00015931 HLA17537 - - - - - - Dr. Nezih Cereb
DRB4*01:52 HG00031358 HLA17537 - - - - - - Dr. Nezih Cereb
DRB4*01:53 HG00013179
HG00013180		 HLA17538 - - - - - - Dr. Nezih Cereb
DRB4*01:54:01N HG00013181 HLA17539 - - - - - - Dr. Nezih Cereb
DRB4*01:54:01N HG00014679 HLA17539 - - - - - - Dr. Nezih Cereb
DRB4*01:54:02N HG00027340 HLA28543 - - - - - - Dr. Nezih Cereb
DRB4*01:55 HG00013182 HLA17540 - - - - - - Dr. Nezih Cereb
DRB4*01:56N HG00013183 HLA17541 - - - - - - Dr. Nezih Cereb
DRB4*01:57N HG00013477 HLA17943 - - - - - - Dr. Nezih Cereb
DRB4*01:57N HG00014621 HLA17943 - - - - - - Dr. Nezih Cereb
DRB4*01:58 HG00013478 HLA17944 - - - - - - Dr. Nezih Cereb
DRB4*01:58 HG00014676 HLA17944 - - - - - - Dr. Nezih Cereb
DRB4*01:58 HG00031356 HLA17944 - - - - - - Dr. Nezih Cereb
DRB4*01:59 HG00013479 HLA17945 - - - - - - Dr. Nezih Cereb
DRB4*01:60 HG00013480 HLA17946 - - - - - - Dr. Nezih Cereb
DRB4*01:60 HG00013829
HG00015298		 HLA17946 - - - - - - Dr. Nezih Cereb
DRB4*01:60 HG00026844 HLA17946 - - - - - - Dr. Nezih Cereb
DRB4*01:60 HG00031357 HLA17946 - - - - - - Dr. Nezih Cereb
DRB4*01:61:01N HG00013482 HLA17947 - - - - - - Dr. Nezih Cereb
DRB4*01:61:02N HG00051229 HLA38076 - - - - - - Dr. Nezih Cereb
DRB4*01:62 81163755 HLA18068 - - - - - - Dr. Daniel Fuerst
DRB4*01:63 81153001 HLA18085 - - - - - - Dr. Daniel Fuerst
DRB4*01:64 HG00013939
HG00013940		 HLA18632 - - - - - - Dr. Nezih Cereb
DRB4*01:65N HG00013785 HLA18684 - - - - - - Dr. Nezih Cereb
DRB4*01:65N HG00018095 HLA18684 - - - - - - Dr. Nezih Cereb
DRB4*01:65N HG00015917 HLA18684 - - - - - - Dr. Nezih Cereb
DRB4*01:66 HG00014675 HLA18792 - - - - - - Dr. Nezih Cereb
DRB4*01:67 HG00014678 HLA18793 - - - - - - Dr. Nezih Cereb
DRB4*01:68 HG00014682 HLA18794 - - - - - - Dr. Nezih Cereb
DRB4*01:69 HG00013553 HLA18820 - - - - - - Dr. Nezih Cereb
DRB4*01:70 HG00017136 HLA19679 - - - - - - Dr. Nezih Cereb
DRB4*01:70 HG00022276 HLA19679 - - - - - - Dr. Nezih Cereb
DRB4*01:70 HG00019608 HLA19679 - - - - - - Dr. Nezih Cereb
DRB4*01:71N HG00017240 HLA19826 - - - - - - Dr. Nezih Cereb
DRB4*01:72:01 HG00016340 HLA20687 - - - - - - Dr. Nezih Cereb
DRB4*01:72:01 HG00022284 HLA20687 - - - - - - Dr. Nezih Cereb
DRB4*01:72:01 HG00022282 HLA20687 - - - - - - Dr. Nezih Cereb
DRB4*01:72:01 HG00025632 HLA20687 - - - - - - Dr. Nezih Cereb
DRB4*01:72:02 23111555279HS27822 HLA39019 - - - - - - Dr. Cathi Murphey
DRB4*01:73 HG00016341 HLA20931 - - - - - - Dr. Nezih Cereb
DRB4*01:73 HG00022269 HLA20931 - - - - - - Dr. Nezih Cereb
DRB4*01:74 HG00016609 HLA20804 - - - - - - Dr. Nezih Cereb
DRB4*01:74 HG00022293 HLA20804 - - - - - - Dr. Nezih Cereb
DRB4*01:74 HG00022287
HG00022300		 HLA20804 - - - - - - Dr. Nezih Cereb
DRB4*01:74 HG00022263 HLA20804 - - - - - - Dr. Nezih Cereb
DRB4*01:75 HG00017497 HLA21033 - - - - - - Dr. Nezih Cereb
DRB4*01:75 HG00018610 HLA21033 - - - - - - Dr. Nezih Cereb
DRB4*01:75 HG00015297
HG00015694
HG00018859
HG00018991
HG00018993
HG00019715		 HLA21033 - - - - - - Dr. Nezih Cereb
DRB4*01:76:01 HG00017505 HLA20820 - - - - - - Dr. Nezih Cereb
DRB4*01:76:01 HG00017573 HLA20820 - - - - - - Dr. Nezih Cereb
DRB4*01:76:02 HG00029514 HLA30928 - - - - - - Dr. Nezih Cereb
DRB4*01:77 HG00018097 HLA20749 - - - - - - Dr. Nezih Cereb
DRB4*01:77 HG00018097 HLA20749 - - - - - - Dr. Nezih Cereb
DRB4*01:78 HG00018098 HLA20625 - - - - - - Dr. Nezih Cereb
DRB4*01:78 HG00022294 HLA20625 - - - - - - Dr. Nezih Cereb
DRB4*01:78 HG00018987
HG00018988
HG00019710		 HLA20625 - - - - - - Dr. Nezih Cereb
DRB4*01:79 HG00018100 HLA20586 - - - - - - Dr. Nezih Cereb
DRB4*01:80N HG00017500
HG00017501		 HLA21562 - - - - - - Dr. Nezih Cereb
DRB4*01:80N HG00017238
HG00017241		 HLA21562 - - - - - - Dr. Nezih Cereb
DRB4*01:81 HG00018847 HLA21227 - - - - - - Dr. Nezih Cereb
DRB4*01:81 HG00022296 HLA21227 - - - - - - Dr. Nezih Cereb
DRB4*01:82 HG00018848 HLA21568 - - - - - - Dr. Nezih Cereb
DRB4*01:83 HG00018849 HLA21459 - - - - - - Dr. Nezih Cereb
DRB4*01:84N HG00018850 HLA21216 - - - - - - Dr. Nezih Cereb
DRB4*01:85 Ua2018tp2849 HLA21905 - - - - - - Dr. Mats Bengtsson
DRB4*01:85 90P223809 HLA21905 - - - - - - Dr. Daniel Fuerst
DRB4*01:86 81192102 HLA21774 - - - - - - Dr. Daniel Fuerst
DRB4*01:87 HG00018612 HLA22622 - - - - - - Dr. Nezih Cereb
DRB4*01:88 HG00018989 HLA22631 - - - - - - Dr. Nezih Cereb
DRB4*01:88 HG00022271
HG00022274		 HLA22631 - - - - - - Dr. Nezih Cereb
DRB4*01:89 HG00018990 HLA22349 - - - - - - Dr. Nezih Cereb
DRB4*01:89 HG00013828 HLA22349 - - - - - - Dr. Nezih Cereb
DRB4*01:90 HG00019610 HLA22202 - - - - - - Dr. Nezih Cereb
DRB4*01:91 HG00019711 HLA22588 - - - - - - Dr. Nezih Cereb
DRB4*01:92 HG00019712 HLA22397 - - - - - - Dr. Nezih Cereb
DRB4*01:93 HG00019713 HLA22252 - - - - - - Dr. Nezih Cereb
DRB4*01:94 HG00022298 HLA23596 - - - - - - Dr. Nezih Cereb
DRB4*01:95 HG00022301 HLA23137 - - - - - - Dr. Nezih Cereb
DRB4*01:96 HG00022289 HLA23132 - - - - - - Dr. Nezih Cereb
DRB4*01:97 HG00022290 HLA23501 - - - - - - Dr. Nezih Cereb
DRB4*01:98 HG00013554 HLA23615 - - - - - - Dr. Nezih Cereb
DRB4*01:99 HG00013825 HLA23389 - - - - - - Dr. Nezih Cereb
DRB4*01:100 HG00013826 HLA23628 - - - - - - Dr. Nezih Cereb
DRB4*01:101 HG00013827 HLA23381 - - - - - - Dr. Nezih Cereb
DRB4*01:102 HG00013830 HLA23321 - - - - - - Dr. Nezih Cereb
DRB4*01:102 114670 HLA23321 - - - - - - Dr. Francisco de Paula Sanchez Gordo
DRB4*01:103 HG00022272 HLA23313 - - - - - - Dr. Nezih Cereb
DRB4*01:104 HG00022273 HLA23054 - - - - - - Dr. Nezih Cereb
DRB4*01:105 HG00022283 HLA23639 - - - - - - Dr. Nezih Cereb
DRB4*01:106 HG00022285 HLA23452 - - - - - - Dr. Nezih Cereb
DRB4*01:106 HG00015916
HG00016612		 HLA23452 - - - - - - Dr. Nezih Cereb
DRB4*01:107 HG00017498 HLA23438 - - - - - - Dr. Nezih Cereb
DRB4*01:107 HG00022502 HLA23438 - - - - - - Dr. Nezih Cereb
DRB4*01:108N HG00014680 HLA24284 - - - - - - Dr. Nezih Cereb
DRB4*01:109 HG00015299 HLA24228 - - - - - - Dr. Nezih Cereb
DRB4*01:110 HG00015692 HLA23912 - - - - - - Dr. Nezih Cereb
DRB4*01:111 HG00016610 HLA24130 - - - - - - Dr. Nezih Cereb
DRB4*01:112 HG00016613 HLA24266 - - - - - - Dr. Nezih Cereb
DRB4*01:112 HG00017509 HLA24266 - - - - - - Dr. Nezih Cereb
DRB4*01:112 HG00031354 HLA24266 - - - - - - Dr. Nezih Cereb
DRB4*01:113N HG00017239
HG00018099
HG00019709		 HLA24119 - - - - - - Dr. Nezih Cereb
DRB4*01:114 HG00017511 HLA23830 - - - - - - Dr. Nezih Cereb
DRB4*01:115N HG00019607 HLA24025 - - - - - - Dr. Nezih Cereb
DRB4*01:116 HG00022487 HLA23858 - - - - - - Dr. Nezih Cereb
DRB4*01:117 HG00018617 HLA24749 - - - - - - Dr. Nezih Cereb
DRB4*01:118 HG00025060 HLA24874 - - - - - - Dr. Nezih Cereb
DRB4*01:118 HG00027345 HLA24874 - - - - - - Dr. Nezih Cereb
DRB4*01:118 HG00050360 HLA24874 - - - - - - Dr. Nezih Cereb
DRB4*01:119 HG00025633 HLA25855 - - - - - - Dr. Nezih Cereb
DRB4*01:119 6123467 HLA25855 - - - - - - Dr. Antonio Balas
DRB4*01:119 114922 HLA25855 - - - - - - Dr. Francisco de Paula Sanchez Gordo
DRB4*01:119 HG00029060 HLA25855 - - - - - - Dr. Nezih Cereb
DRB4*01:120 HG00025634 HLA25994 - - - - - - Dr. Nezih Cereb
DRB4*01:121N HG00025635 HLA25805 - - - - - - Dr. Nezih Cereb
DRB4*01:121N HG00027349 HLA25805 - - - - - - Dr. Nezih Cereb
DRB4*01:122 HG00025853 HLA26299 - - - - - - Dr. Nezih Cereb
DRB4*01:123 84028210 HLA26580 - - - - - - Dr. Daniel Fuerst
DRB4*01:124 UOU0016
UOU0054
UOU0056		 HLA26722 - - - - - - Dr. Eszter Lazar-Molnar
DRB4*01:125 c0546109 HLA26946 - - - - - - Dr. Francisco de Paula Sanchez Gordo
DRB4*01:126 HG00027356 HLA28536 - - - - - - Dr. Nezih Cereb
DRB4*01:127 HG00027343
HG00027352		 HLA28537 - - - - - - Dr. Nezih Cereb
DRB4*01:127 HG00029078 HLA28537 - - - - - - Dr. Nezih Cereb
DRB4*01:127 HG00029066
HG00029076
HG00029084		 HLA28537 - - - - - - Dr. Nezih Cereb
DRB4*01:128N HG00027344 HLA28538 - - - - - - Dr. Nezih Cereb
DRB4*01:128N HG00029511 HLA28538 - - - - - - Dr. Nezih Cereb
DRB4*01:128N HG00050362 HLA28538 - - - - - - Dr. Nezih Cereb
DRB4*01:129 HG00027357 HLA28539 - - - - - - Dr. Nezih Cereb
DRB4*01:130 HG00027358 HLA28540 - - - - - - Dr. Nezih Cereb
DRB4*01:131 HG00027347 HLA28541 - - - - - - Dr. Nezih Cereb
DRB4*01:132 HG00027348 HLA28542 - - - - - - Dr. Nezih Cereb
DRB4*01:133 HG00027355 HLA28545 - - - - - - Dr. Nezih Cereb
DRB4*01:134 20211439 HLA28680 - - - - - - Dr. Daniel Fuerst
DRB4*01:135 HG00026845 HLA28874 - - - - - - Dr. Nezih Cereb
DRB4*01:136 HG00026843 HLA28875 - - - - - - Dr. Nezih Cereb
DRB4*01:136 HG00027350
HG00027353		 HLA28875 - - - - - - Dr. Nezih Cereb
DRB4*01:137 HG00026846 HLA28876 - - - - - - Dr. Nezih Cereb
DRB4*01:138 HG00026386 HLA29155 - - - - - - Dr. Nezih Cereb
DRB4*01:138 HG00027359
HG00029061
HG00029065
HG00029074		 HLA29155 - - - - - - Dr. Nezih Cereb
DRB4*01:138 HG00029068 HLA29155 - - - - - - Dr. Nezih Cereb
DRB4*01:138 HG00029513 HLA29155 - - - - - - Dr. Nezih Cereb
DRB4*01:138 HG00031521 HLA29155 - - - - - - Dr. Nezih Cereb
DRB4*01:138 HG00049819 HLA29155 - - - - - - Dr. Nezih Cereb
DRB4*01:139 HG00029072 HLA29839 - - - - - - Dr. Nezih Cereb
DRB4*01:140 HG00029070 HLA30269 - - - - - - Dr. Nezih Cereb
DRB4*01:141 HG00029077 HLA30386 - - - - - - Dr. Nezih Cereb
DRB4*01:142 HG00029083 HLA30343 - - - - - - Dr. Nezih Cereb
DRB4*01:143 HG00029082 HLA30378 - - - - - - Dr. Nezih Cereb
DRB4*01:143 HG00050363 HLA30378 - - - - - - Dr. Nezih Cereb
DRB4*01:144 HG00029075 HLA30430 - - - - - - Dr. Nezih Cereb
DRB4*01:145N HG00029062
HG00029086		 HLA30677 - - - - - - Dr. Nezih Cereb
DRB4*01:146 21203211 HLA30601 - - - - - - Dr. Daniel Fuerst
DRB4*01:147 HG00029508 HLA31372 - - - - - - Dr. Nezih Cereb
DRB4*01:148 HG00029515 HLA31395 - - - - - - Dr. Nezih Cereb
DRB4*01:149N HG00029714 HLA31451 - - - - - - Dr. Nezih Cereb
DRB4*01:150 HG00030143 HLA31977 - - - - - - Dr. Nezih Cereb
DRB4*01:151 2110307078 HLA32287 Cargou M, Andreani M, Troiano M, Wojciechowski E, Visentin J Characterization of the novel HLA-DRB4*01:151 allele by sequencing-based typing. HLA 2022 99 64-66 Dr. Jonathan Visentin
DRB4*01:152 KUHSC-24-21 HLA32168 Ameen R, Geo JA, Thomas J Identification of a novel HLA-DRB4 allele, HLA-DRB4*01:152 in a Kuwaiti family. HLA 2022 100 546-547 Dr. Reem Ameen
DRB4*01:153 HG00031359 HLA32726 - - - - - - Dr. Nezih Cereb
DRB4*01:154 HG00031360 HLA32911 - - - - - - Dr. Nezih Cereb
DRB4*01:155 2022S94 HLA34347 - - - - - - Dr. Alexandre Walencik
DRB4*01:156 22S140 HLA34373 - - - - - - Dr. Alexandre Walencik
DRB4*01:157 HG00031980 HLA34262 - - - - - - Dr. Nezih Cereb
DRB4*01:158 HG00027346 HLA35307 - - - - - - Dr. Nezih Cereb
DRB4*01:159N HG00015691 HLA34925 - - - - - - Dr. Nezih Cereb
DRB4*01:160 HG00049910
HG00050164		 HLA34958 - - - - - - Dr. Nezih Cereb
DRB4*01:161 HG00050076 HLA34898 - - - - - - Dr. Nezih Cereb
DRB4*01:162N YUMC 006 HLA35013 Kim SW, Hong JY, Jung S, Lee J, Park Y Identification of the novel HLA-DRB4*01:162N allele using next-generation sequencing. HLA 2022 100 659-660 Dr. Yoonhee Park
DRB4*01:163 HG00050288 HLA35953 - - - - - - Dr. Nezih Cereb
DRB4*01:164 5997073-264317 HLA36154 - - - - - - Dr. Neifi Hassan Saloum Deghaide
DRB4*01:165N HILAPL-001 HLA36512 Hu Q, Burns C, McKinley C, Christian D, Berka N Identification of a novel allele with a frameshift mutation, HLA-DRB4*01:165N, using next-generation sequencing. HLA 2023 101 699-700 Dr. Qian Hu
DRB4*01:166 HG00050547 HLA36453 - - - - - - Dr. Nezih Cereb
DRB4*01:167 5996042-263676 HLA36603 - - - - - - Dr. Neifi Hassan Saloum Deghaide
DRB4*01:168 22S181 HLA36964 - - - - - - Dr. Alexandre Walencik
DRB4*01:169 TPH22-49-F2 HLA37076 - - - - - - Dr. Antonio Nieto
DRB4*01:170 HG00050515 HLA37332 - - - - - - Dr. Nezih Cereb
DRB4*01:171 HG00050948 HLA37293 - - - - - - Dr. Nezih Cereb
DRB4*01:172 81279727 HLA37189 - - - - - - Dr. Daniel Fuerst
DRB4*01:173 23002272 HLA38034 - - - - - - Dr. Robert Bray
DRB4*01:174 HG00051230 HLA38073 - - - - - - Dr. Nezih Cereb
DRB4*01:175 521/23 HLA38471 - - - - - - Mr. Rafael Oliveira
DRB4*01:176 139459821 HLA38466 - - - - - - Dr. Pascal Pedini
DRB4*01:177 HG00051309 HLA38352 - - - - - - Dr. Nezih Cereb
DRB4*01:178 HG00051424 HLA38705 - - - - - - Dr. Nezih Cereb
DRB4*01:179 SZ-213 HLA39346 - - - - - - Dr. Hong-Yan Zou
DRB4*01:180 24202090 HLA39729 - - - - - - Dr. Daniel Fuerst
DRB4*01:181 HG00051934 HLA40065 - - - - - - Dr. Nezih Cereb
DRB4*02:01N GN016 HLA00913 Robbins F, Hurley CK, Tang T, Yao H, Lin YS, Wade J, Goeken N, Hartzman RJ Diversity associated with the second expressed HLA-DRB locus in the human population Immunogenetics 1997 46 104-110 Dr. Fu-Meei Robbins
DRB4*02:01N GN016 HLA00913 - - - - - - Prof. Carolyn K Hurley
DRB4*03:01N GN017 HLA00914 Robbins F, Hurley CK, Tang T, Yao H, Lin YS, Wade J, Goeken N, Hartzman RJ Diversity associated with the second expressed HLA-DRB locus in the human population Immunogenetics 1997 46 104-110 Dr. Fu-Meei Robbins
DRB4*03:01N GN017
GN020
GN076
GN092		 HLA00914 Steiner NK, Hou L, Hurley CK Characterizing allele with large deletions using region specific extraction Human Immunology 2018 79 491-493 Prof. Carolyn K Hurley